Figure 1:

Figure 2:

Figure 3:

Figure 4:

Figure 5:

Data matrix for cluster analysis of the species of Pelodera ‘strongyloides’ groupa_
| Characters | 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 10 | 11 | 12 | 13 | 14 | 15 | 16 | 17 | 18 | 19 | 20 | 21 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| P. cutanea | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 1 | 1 | 1 | 1 |
| P. nidicolis | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 2 | 1 | 0 | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
| P. orbitalis | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 |
| P. strongyloides | 0 | 2 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 2 | 1 | 0 | 2 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 |
| P. strongyloides ex Lucanus ibericus | 0 | 2 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 |
| P. termitis | 0 | 0 | 0 | 0 | 1 | 1 | ? | 0 | 1 | 1 | 0 | 1 | 1 | 0 | 1 | 1 | 2 | 0 | 0 | 0 | 0 |
| P. scrofulata | 1 | 1 | 1 | 1 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 1 |
Ratios and measurements (in µm) of heat relaxed specimens of Pelodera strongyloides collected from Lucanus ibericus in Georgia_
| Character | Females | Males | 3rd juvenile stage | P. strongyloides (Schneider, 1866) |
|---|---|---|---|---|
| n | 20 | 20 | 22 | 11 |
| Body length | 1566 ± 120 (1353-1867) | 1280 ± 100 (1080-1494) | 663 ± 51 (566-772) | 1629 (1142-2149) |
| Body width | 96 ± 10 (75-112 | 62 ± 4.8 (55-70) | 30 ± 2.4 (25-35) | 107 (80-132) |
| Lip region diam | 19 ± 1.6 (16-22 | 17 ± 1.5 (15-20) | – | – |
| Stoma length | 31 ± 1 (30-32) | 29 ± 2.2 (25-32) | 24 ± 1.7 (22-30) | 33 (30-38) |
| Stoma diam | 5.1 ± 0.4 (5.0-6.2) | 4.7 ± 0.4 (3.7-5.0) | 2.6 ± 0.4 (2.5-3.7) | – |
| Anterior end to nerve ring | 178 ± 6 (172-197) | 155 ± 10 (137-175) | 102 ± 6 (92-120) | – |
| Stoma length as % pharynx length | 12.2 ± 0.4 (11-13) | 13 ± 1.2 (11-15) | 16 ± 0.8 (14-17) | – |
| Pharynx length | 252 ± 5 (237-257) | 219 ± 17 (192-250) | 152 ± 6 (142-177) | – |
| Corpus length | 144 ± 7 (135-162) | 122 ± 10 (107-137) | 87 ± 2 (80-92) | – |
| Corpus as % pharynx length | 56 ± 2.3 (53-63) | 55 ± 2 (50-59) | 57 ± 2.5 (52-61) | – |
| Median bulb (MB) diam | 27 ± 2.1 (22-30) | 30 ± 4.9 (20-37) | 11 ± 0.8 (10-12) | – |
| Terminal bulb (TB) diam | 39 ± 4.1 (32-47) | 34 ± 2.7 (32-40) | 15 ± 0.6 (12-17) | 36-50 |
| MB diam. as % TB diam | 77 ± 9.5 (54-88) | 79 ± 6.8 (62-93) | – | – |
| Anterior end to excretory pore | 236 ± 14 (212-270) | 205 ± 13 (164-227) | 130 ± 10 (110-150) | 235 (183-295) |
| Ex. pore posit. as % phar. length | 93 ± 5 (83-105) | 93 ± 8 (75-109) | 85 ± 6.7 (75-98) | – |
| Tail length | 69 ± 11 (52-95) | 53 ± 5 (45-62) | 72 ± 4 (65-82) | – |
| Anal (cloacal) body width (ABW) | 40 ± 3 (32-47) | 37 ± 3 (32-42) | 19 ± 1.8 (16-22) | – |
| Gonad lengtha | 1285 ± 198 (995-1658) | 956 ± 87 (747-1268) | – | 890 (653-1157) |
| Gonad length as % body length | 82 ± 15 (61-107) | 72 ± 5 (66-82) | – | 46-63 |
| Gonad length. as % intestine leng.b | 108 ± 23 (72-143) | 93 ± 11 (69-110 ) | – | – |
| Anterior gonad branch length | 800 ± 104 (555-928) | – | – | – |
| Posterior branch length | 546 ± 128 (290-794) | – | – | – |
| Testis flexure length | – | 201 ± 49 (123-260) | – | – |
| Ant. flexure as % of ant. branch | 33 ± 3 (26-42) | 17 ± 4 (13-24) | – | – |
| Post. flex. as % of post. branch | 27 ± 6 (13-34) | – | – | – |
| Sperm diameter | n.d. | 5 ± 0.6 3. (7-7.5) | – | – |
| Egg length | 60 ± 3.4 (55-67) | – | – | – |
| Egg diam | 41 ± 4 (35-47) | – | – | – |
| Rectum length | 38 ± 3.2 (30-42) | n.d. | 24 ± 1.5 (22-25) | – |
| Rectum length/ABW | 0.9 ± 0.1 (0.8-1.3) | n.d. | 0.9 ± 0.1 (7-1.2) | – |
| Anus to phasmid distance | 23 ± 1.8 (20-27) | n.d. | – | – |
| Anus to phasmid distance/ABW | 0.6 ± 0.05 (0.4-0.7) | n.d. | – | – |
| Posit. phasmid as % tail length | 35 ± 6 (27-48) | – | – | 32-61 |
| Spicule length | – | 66 ± 3 (60-72) | – | 55-82 |
| Gubernaculum length | – | 44 ± 2.2 (40-50) | – | – |
| Gubern. leng. as % spic. length | – | 66 ± 4.5( 60-78) | – | – |
| a | 16.3 ± 2 (13.5-20.3) | 20.7 ± 1 (19-23.1) | 22.3 ± 1.5 (19.3-25.8) | 15.1 (12-18.7) |
| b | 6.1 ± 0.4 (5.4-7) | 5.8 ± 0.5 (5.1-7) | 4.2 ± 0.2 (3.7-4.7) | 6.2 (4.9-7.6) |
| c | 23 ± 3.4 (17.6-32.2) | 24.3 ± 2.8 (20.5-30.4) | 9.1 ± 0.5 (8-10) | 33 (26.3-37.9) |
| c′ (= tail length/ABW) | 1.6 ± 0.6 (1.3-2.2) | 1.3 ± 0.1 (1.2-1.7) | 3.7 ± 0.3 (3.2-4.5) | – |
| V (vulva posit. in % body length) | 57 ± 1.9 (51.8-59.1) | – | – | 58 (55-64) |