Skip to main content
Have a personal or library account? Click to login
Petiole anatomical study in some taxa of the genus Alcea (Malvaceae) from Iran Cover

Petiole anatomical study in some taxa of the genus Alcea (Malvaceae) from Iran

Open Access
|Nov 2026
Read online

References

  1. Alefeld, F. G. C. (1862). Über die Malveen. Österreichische botanische Zeitschrift, 22, 247–261.
  2. Azizian, D., & Cutler, D. F. (1982). Anatomical, cytological and phytochemical studies on Phlomis L. and Eremostachys Bunge (Labiatae). Botanical Journal of the Linnean Society, 85, 249–281.
  3. Baker, E.G. (1890). Synopsis of genera and species of Malvae. Journal of Botany, 28, 140–145, 207–209.
  4. Boissier, P. E. (1867). Flora orientalis, vol. 1. Genevae et Baseileae.
  5. Cabi, E., Başer, B., Uzunhisarcikli, M. E., & Yavru, A. (2009). Pollen morphology of Alcea L. and Althaea L. genera (Malvaceae) in Turkey. Feddes Repertorium, 120(7–8), 405–418. http://doi:10.1002/FEDR.200911119
  6. Candolle, A. P. de. (1828). Prodromus sytematis naturalis regni vegetabilis, vol. 3 (pp. 207–296). Sumptibus Sociorum Treuttel et Würtz.
  7. Escobar Garcia, P., Pakravan, M., Schönswetter, P., Fuertes Aguilar, J., & Schneeweiss, G.M. (2012). Phylogenetic relationships in the species-rich Irano-Turanian genus Alcea (Malvaceae). Taxon, 61(2), 324–332. http://doi:10.1002/tax.612004.
  8. Iljin, M.M. (1949). Malvaceae. In: V. L., Komarov, B. K., Shishkin, & E. G., Bobrov (Eds.), Flora SSSR, vol. 15 (pp. 21–137). Botanical Institute of the Academy of Sciences of the USSR, Leningrad.
  9. Linnaeus, C. (1742). Genera Plantarum. C. Wishoff et G. J. Wishoff, Leiden.
  10. Marasek-Ciolakowska, A., Soika, G., Warabieda, W., Kowalska, U., & Rybczynski, D. (2021). Investigation on the relationship between morphological and anatomical characters of Savoy cabbage and kale leaves and investigation by cabbage whitefly (Aleyrodes proletella L.). Agronomy, 11(2), 275.
  11. Metcalfe, C. R., & Chalk, L. (1979). Anatomy of the Dicotyledons. vol. 1 (pp. 450–464). Clarendron Press.
  12. Mohammad Ebrahim, Z., Hassan, S., Elazab, H., M., & Badawi, A. (2018). Cladistic analysis of some taxa in Malvaceae s. l. “Core Malvales” based on anatomical characteristics. The Egyptian Journal of Experimental Biology (Bot.), 14(1), 87–105. http://doi:5455/egyjebb.20180210103057
  13. Moon, H., Hong, S., Smets, E., & Huysmans, S. (2009). Phylogenetic significance of leaf micromorphology and anatomy in the tribe Mentheae (Nepetoideae: Lamiaceae). Botanical Journal of the Linnean Society, 160(2), 211–231. http://doi:10.1111/j.1095-8339.2009.00979.x
  14. Nobarinezhad, M. H., Pakravan, M., & Pahlevani, A. (2018). A biosystematic study of Euphorbia subgenus Chamaesyce (Euphorbiaceae) in Iran. Phytotaxa, 360, 179–200. https://doi:org/10.11646/phytotaxa.360.3.1
  15. Özbek, F., & Uzunhisarcıkli, M. E. (2023). Taxonomic significance of seed macro-micromorphology of Turkish Alcea L. (Malvaceae) through light microscopy and scanning electron microscopy. Microscopy Research and Technique, 86(12), 1551–1567. https://doi.org/10.1002/jemt.24385
  16. Ozkan, A. M., & Uzunhisarcıkli, M. E. (2007). Stem and Leaf Anatomy of Althaea L. (Malvaceae) Species Growing in Turkey. Hacettepe University Journal of the Faculty of Pharmacy, 28(2), 133–148.
  17. Pakravan, M. (2001). Biosystematic study of the genus Alcea L. (Malvaceae) in Iran. Dissertation, University of Tehran, Tehran.
  18. Pakravan, M. (2008). Malvaceae. In M., Assadi, et al. (Eds), Flora of Iran, No. 58. RIFR, Tehran.
  19. Pakravan, M. (2025). Seed protein analysis as a tool for taxonomy of Alcea (Malvaceae) in Iran. Plant, Algae, and Environment, 9(3), 94–106.
  20. Riedl, I. (1976). Alcea. 41–80. In: K. H., Rechinger (Ed), Flora Iranica, 120. Akademische Druck- und Verlagsanstalt.
  21. Schweingruber, F. H., Riha, P., & Doležal, J. (2014). Variation in stem anatomical characteristics of Campanuloideae species in relation to evolutionary history and ecological preferences. Plos One, 9, e88199. https://doi:10.1371/journal.pone.0088199
  22. Shaheen, N. I., Khan, M. A., Yasmin, G., Hayat, M. Q. Munsif, S. H., & Ahmad, K. H. (2010). Foliar epidermal anatomy and pollen morphology of the genera Alcea and Althaea (Malvaceae) from Pakistan. International Journal of Agriculture and Biology, 12(3), 329–324.
  23. Silva, K. R., Romero, R., & Simăo, D. G. (2018). Leaf characters of Lavoisiera microlicia and Trembleya (Melastomataceae) and their implications for taxonomy. Feddes Repertorium, 129(2), 123–136. https://doi.org/10.1002/fedr.201700019
  24. Talip, N., Cutler, D. F., Ahmad Puad, A. S., Ismail, B. S., Ruzi, A. R., & Ahmad Juhari, A. A. (2017). Diagnostic and systematic significance of petiole anatomy in the identification of Hopea species (Dipterocarpaceae). South African Journal of Botany, 111, 111–125
  25. Thacker, K. D., Gavade, S. K., Lekhak, M. M., Gondaliya, A. D., & Rajput, K. S. (2021). Comparison of petiole anatomy in Flemingia and its potential for delimitation of species. Flora, 278, 151790. https://doi.org/10.1016/j.flora.2021.151790
  26. Thiers, B. M. (2025). Continuously updated. Index herbariorum: A directory of public herbarium and associated staff. New yourk Botanical Garden’s Virtual Herbarium. https://sweetgum.nybg.org/science/ih/
  27. Tournefort, J. (1700). Institutiones rei herbariae-E Typographia Regia, pp. 105–107. Paris.
  28. Willdenow, C. V. (1800). Species plantarum, ed. 4, 3. Impensis GC Nauk, Berolinum [Berlin].
  29. Zohary, M. (1963a). Taxonomical studies in Alcea of South-Western Asia. Part I. Bulletin of the Research Council of Israel, 11(D4), 210–229.
  30. Zohary, M. (1963b). Taxonomical studies in Alcea of South-Western Asia. Part II. Israel Journal of Botany, 121, 1–26.
DOI: https://doi.org/10.3986/hacq-2026-0003 | Journal eISSN: 1854-9829 | Journal ISSN: 1581-4661
Language: English, Slovenian
Page range: 355 - 364
Submitted on: Apr 12, 2025
Accepted on: Feb 14, 2026
Published on: Nov 4, 2026
Published by: Slovenian Academy of Sciences and Arts
In partnership with: Paradigm Publishing Services

© 2026 Maneezheh Pakravan, Azam Javanbakht, published by Slovenian Academy of Sciences and Arts
This work is licensed under the Creative Commons Attribution 4.0 License.