
Figure 1.
Location of the study plots
Table 1.
Characteristics of habitat conditions in the central and northern Poland (in 2007–2017)
| Forest District | Average annual temperature [°C] | Annual sum of precipitation [mm] | Soil type | Parent material origin | Type of forest habitat | Dominant tree species |
|---|---|---|---|---|---|---|
| Białowieża | 7.0 | 550 | BRt | Qp | BMśw | So |
| Chojnów | 8.0 | <550 | RDb | QZp | BMśw | So |
| Gdańsk | 7.5 | 650 | BRKt | Qp | Lśw | Bk |
| Krotoszyn | 8.0 | <550 | OGw | Qg | Lśw | Db |
| Krucz | 8.0 | <550 | B | QEp/QZp | Bśw | So |
| Łąck | 8.0 | <550 | RDbr | QFp | LMśw | Db |
| Strzałowo | 7.5 | 650 | Pbr | QFp | BMśw | So |
| Suwałki | 6.5 | 550 | BRg | Qg | Lśw | Św |
1 Soil type: BRt – Eutric Cambisol, RDb – Brunic Arenosol (Protospodic), BRKt – Dystric Cambisol, OGw – Stagnosol, B – Haplic Podzol, RDbr – Brunic Arenosol, Pbr – Haplic Luvisol, BRg – Gleyic Cambisol; Parent material origin: Qp – glacial sand, QZp – fluvioglacial sand (sandr), Qg – glacial till, QEp/QZp – aeolian sand over fluvioglacial sand (sandr), QFp – fluvioglacial sand; Habitat type of the forest: BMśw – mixed fresh coniferous forest, Lśw – fresh broadleaved forest, Bśw – fresh coniferous forest, LMśw – fresh mixed broadleaved forest; Dominant tree species: So – Scots pine, Bk – European beech, Db – Oak, Św – Norway spruce.

Figure 2.
Particle size distribution on the study plots
Table 2.
Average of pH values and TOC and TN contents in forest soils in central and northern Poland (2007–2017)
| Forest District | Layer (cm) | Year | pHCaCl2 | TOC [g·kg-1] | TN [g·kg-1] | C :N |
|---|---|---|---|---|---|---|
| 1 | 2 | 3 | 4 | 5 | 6 | 7 |
| Chojnów | O | 2007 | 3.20 | 313.90 | 13.75 | 22.83 |
| 2017 | 3.19 | 390.38 | 17.16 | 22.75 | ||
| mean±SD | 3.19±0.10 | 325.48±48.67 | 15.45±2.41 | 22.79±1.14 | ||
| M | 2007 | 3.84a’ | 7.53a’ | 0.42a’ | 16.97a’ | |
| 2017 | 3.71a’ | 9.04a’ | 0.46a’ | 18.54a’ | ||
| mean±SD | 3.78±0.40 | 8.53±4.94 | 0,44±0.18 | 17.74±3.52 | ||
| Łąck | O | 2007 | 4.50 | 218.50 | 11.19 | 19.53 |
| 2017 | 4.50 | 268.87 | 12.68 | 21.19 | ||
| mean±SD | 4.50±0.33 | 239.48±26.21 | 11.93±1.05 | 20.36±0.83 | ||
| M | 2007 | 3.88a’ | 6.10a’ | 0.38a’ | 12.34a’ | |
| 2017 | 4.04a’ | 12.51a’ | 0.64a’ | 17.60a’ | ||
| mean±SD | 3.99±0.28 | 9.84±9.82 | 0.51±0.49 | 16.66±4.40 | ||
| Krotoszyn | O | 2007 | 3.20 | 305.80 | 16.19 | 18.89 |
| 2017 | 3.20 | 369.48 | 20.91 | 17.67 | ||
| mean±SD | 3.20±0.01 | 353.73±34.55 | 18.55±1.05 | 17.70±6.40 | ||
| M | 2007 | 3.52a’ | 23.38a’ | 1.32a’ | 15.97a’ | |
| 2017 | 3.46a’ | 19.71a’ | 1.00a’ | 17.04a’ | ||
| mean±SD | 3.49±0.36 | 20.08±8.73 | 1.16±0.96 | 17.00±0.98 | ||
| Krucz | O | 2007 | 2.70 | 357.60 | 11.84 | 30.20 |
| 2017 | 2.71 | 399.61 | 13.66 | 29.24 | ||
| mean±SD | 2.70±0.07 | 315.90±90.32 | 12.70±1.29 | 29.54±0.47 | ||
| M | 2007 | 3.81a’ | 7.18a’ | 0.39a’ | 16.82a’ | |
| 2017 | 4.19a’ | 7.09a’ | 0.36a.’ | 16.65a’ | ||
| mean±SD | 3.92±0.43 | 7.11±0.70 | 0.38±0.20 | 16.51a’0.07 | ||
| Suwałki | O | 2007 | 4.10 | 154.00 | 8.07 | 19.08 |
| 2017 | 4.11 | 166.17 | 8.81 | 18.86 | ||
| mean±SD | 4.00±0.14 | 162.99±9.72 | 8.40±0.52 | 18.69±0.83 | ||
| M | 2007 | 3.73a’ | 12.79a’ | 1.02a’ | 11.24a’ | |
| 2017 | 3.94a’ | 14.31a’ | 0.99a’ | 14.59a’ | ||
| mean±SD | 3.80±0.29 | 13.90±9.32 | 1.08±0.62 | 13.55±3.04 | ||
| Białowieża | O | 2007 | 3.60 | 289.20 | 16.98 | 22.81 |
| 2017 | 3.59 | 323.39 | 13.92 | 23.22 | ||
| mean±SD | 3.60±0.21 | 326.53±31.83 | 15.45±2.16 | 22.63±0.56 | ||
| M | 2007 | 3.99a’ | 12.51a’ | 0.42a’ | 15.47a’ | |
| 2017 | 4.10a’ | 17.01a’ | 0.46a’ | 16.34a’ | ||
| mean±SD | 4.07±0.47 | 14.67±12.02 | 0.46±0.20 | 16.19±1.82 | ||
| Strzałowo | O | 2007 | 4.30 | 56.92 | 3.10 | 18.36 |
| 2017 | 4.31 | 162.87 | 6.80 | 23.95 | ||
| mean±SD | 4.30±0.01 | 114.28±57.30 | 4.95±2.61 | 21.04±2.29 | ||
| M | 2007 | 4.51a’ | 8.25a’ | 0,61a’ | 12.38a’ | |
| 2017 | 4.60a’ | 9.74a’ | 0.65a’ | 14.40a’ | ||
| mean±SD | 4.58±0.22 | 9.18±5.82 | 0,63±0,40 | 13.02±1.83 | ||
| Gdańsk | O | 2007 | 4.50 | 303.30 | 16.07 | 18.87 |
| 2017 | 4.51 | 358.88 | 14.54 | 24.68 | ||
| mean±SD | 4.50±0.01 | 326.48±28.91 | 15,30±1.08 | 21.78±2.90 | ||
| M | 2007 | 3.79a’ | 8.51a’ | 0,58a’ | 13.91a’ | |
| 2017 | 3.94a’ | 8.20a’ | 0.51a’ | 14.86a’ | ||
| mean±SD | 3.84±0.30 | 8.36±4.50 | 0,55±0,26 | 14.23±3.10 |

Figure 3.
Selected chemical parameters of forest soils in central and northern Poland in 2007 and 2017 against the background of: A-B) – location of the study areas, C-D) – forest habitat type; E-F) – soil type (WRB)
Table 3.
Selected chemical indicators of the fertility status of forest soils of the central and northern Poland (2007–2017)
| Forest District | Layer | Year | SBC (cmol·kg-1) | CEC (cmol·kg-1) | BS (%) | SBC : Al | Ca : Al |
|---|---|---|---|---|---|---|---|
| 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 |
| Chojnów | O | 2007 | 16.11a | 21.58a | 74.65a | 11.21b | 8.52a |
| 2017 | 16.54a | 20.82a | 79.42a | 9.13b | 6.71a | ||
| mean±SD | 16.32±0.30 | 21.20±0.53 | 77.04±3.37 | 10.1±1.47 | 7.62±1.28 | ||
| M | 2007 | 0.18a’ | 1.34a’ | 13.07a’ | 0.18a’ | 0.11a’ | |
| 2017 | 0.22a’ | 2.09a’ | 12.08a’ | 0.14a’ | 0.08a’ | ||
| mean±SD | 0.20±0.11 | 1.70±0.96 | 12.58±3.87 | 0.16±0.06 | 0.10±0.05 | ||
| Łąck | O | 2007 | 28.95a | 31.03a | 93.30a | 298.90a | 27.60a |
| 2017 | 23.30a | 26.40a | 88.25a | 44.65b | 33.67a | ||
| mean±SD | 26.98±4.00 | 29.63±3.27 | 90.77±3.56 | 116.77±172.71 | 30.63±4.29 | ||
| M | 2007 | 0.28a’ | 1.09a’ | 21.16a’ | 0.30a’ | 0.20’ | |
| 2017 | 0.27a’ | 2.20a’ | 13.68a’ | 0.16a’ | 0.09’ | ||
| mean±SD | 0.27±0.27 | 1.64±1.21 | 17.42±9.01 | 0.23±0.22 | 0.14±0.15 | ||
| Krotoszyn | O | 2007 | 9.18a | 18.68b | 50.80b | 1.92a | 1.24a |
| 2017 | 29.82b | 38.08a | 78.32a | 6.30a | 1.00a | ||
| mean±SD | 19.50±14.6 | 28.07±14.15 | 64.56±19.46 | 4.10±3.09 | 1.22±0.17 | ||
| M | 2007 | 1.58a’ | 5.46a’ | 27.30a’ | 0.57a’ | 0.34a’ | |
| 2017 | 2.67a’ | 7.09a’ | 29.03a’ | 1.33a’ | 0.86a’ | ||
| mean±SD | 2.13±2.83 | 6.27±2.49 | 28.17±23.76 | 0.95±1.75 | 0.60±1.18 | ||
| Krucz | O | 2007 | 6.04a | 15.95a | 37.87b | 2.18a | 1.64a |
| 2017 | 14.57b | 22.15a | 65.76a | 9.10b | 6.77b | ||
| mean±SD | 10.30±6.03 | 19.05±4.38 | 51.81±19.72 | 5.64±4.90 | 4.21±3.62 | ||
| M | 2007 | 0.11a’ | 1.24a’ | 10.67a’ | 0.12a’ | 0.07a’ | |
| 2017 | 0.15a’ | 1.59a’ | 12.73a’ | 0.21a’ | 0.12a’ | ||
| mean±SD | 0.13±0.07 | 1.42±0.94 | 11.70±8.32 | 0.16±0.17 | 0.10±0.11 | ||
| Suwałki | O | 2007 | 12.45a | 14.92a | 83.45a | 25.79a | 21.87a |
| 2017 | 17.17a | 19.38a | 88.59a | 18.75a | 14.39a | ||
| mean±SD | 14.81±3.33 | 17.15±3.15 | 86.02±3.64 | 22.27±5.00 | 18.11±5.32 | ||
| M | 2007 | 0.82a’ | 3.77a’ | 23.36a’ | 0.66a’ | 0.52a’ | |
| 2017 | 4.72a’ | 7.80a” | 36.00a’ | 5.22b’ | 4.22b’ | ||
| mean±SD | 2.10±2.75 | 4.89±3.59 | 26.15±17.14 | 1.79±3.22 | 2.37±3.11 | ||
| Białowieża | O | 2007 | 17.14a | 21.58a | 79.42a | 24.37a | 21.07a |
| 2017 | 7.20b | 11.50b | 62.60a | 4.08b | 3.38b | ||
| mean±SD | 12.17±7.03 | 16.54±7.13 | 71.01±11.90 | 14.24±14.34 | 12.23±12.50 | ||
| M | 2007 | 0.57a’ | 2.61a’ | 28.30a’ | 0.75a’ | 0.64a’ | |
| 2017 | 1.68a’ | 3.66a’ | 51.71b’ | 10.05b’ | 9.62b’ | ||
| mean±SD | 1.02±0.54 | 3.09±2.37 | 37.18±2.37 | 1.11±0.89 | 5.13±1.09 | ||
| Strzałowo | O | 2007 | 7.85a | 8.75a | 89.71a | 22.94b’ | 18.82a |
| 2017 | 16.83b | 21.05b | 79.95a | 7.95a’ | 6.25a | ||
| mean±SD | 12.34±6.35 | 14.90±8.70 | 84.83±6.90 | 15.44±10.60 | 12.54±8.89 | ||
| M | 2007 | 1.74a’ | 2.18a’ | 84.18a’ | 11.21a’ | 10.01a’ | |
| 2017 | 1.67a’ | 2.71a’ | 63.29a’ | 2.49b’ | 2.02b’ | ||
| mean±SD | 1.70±0.55 | 2.44±0.95 | 73.73±18.10 | 6.85±9.45 | 6.02±8.53 | ||
| Gdańsk | O | 2007 | 20.54a | 23.02a | 89.23a | 25.12a | 18.94a |
| 2017 | 27.32a | 32.18a | 84.90a | 20.23a | 15.30a | ||
| mean±SD | 23.93±4.79 | 27.60±6.48 | 87.06±3.06 | 23.51±3.52 | 17.12±2.57 | ||
| M | 2007 | 0.21a’ | 1.34a’ | 15.12a’ | 0.24a’ | 0.14a’ | |
| 2017 | 0.34a’ | 1.71a’ | 21.49a’ | 0.29a’ | 0.14a’ | ||
| mean±SD | 0.23±0.16 | 1.52±0.45 | 18.30±8.75 | 0.26±0.14 | 0.14±0.11 |
1 Layer: O – organic, M – mineral (0–80 cm deph) – arithmetic mean value; SBC – sum of basic cations; CEC – cation exchange capacity; BS% – alkaline saturation; CBC : Al ratio; BS – base saturation, CEC – cation exchange capacity, SBC – sum of base cations, SBC : Al ratio (the ratio of the sum of alkali cations to aluminum).
Table 4.
Non-parametric Spearman correlation between the properties of forest soils in Poland
| Soil type | Layer | Sand | Silt | Clay | Texture | pH | TOC | TN | C :N | SBC | SBC : Al. | CEC | BS | Ca : Al. | |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| TSL | – | – | –0.361 (*) | – | 0.343 (*) | – | – | – | – | – | – | – | – | – | – |
| Tree species | 0.539 (***) | – | – | – | – | –0.367 (*) | – | – | – | – | – | – | – | –0.466(**) | – |
| Layer | – | – | – | – | – | – | 0.743 (***) | –0.618 (***) | –0.637 (***) | –0.668 | – | 0.338 (*) | – | – | – |
| Sand | – | – | – | –0.913 | –0.775 (***) | –0.557 (***) | – | –0.333 (*) | –0.396 (*) | – | –0.702 (***) | – | –0.853 (***) | – | – |
| Silt | – | – | –0.913 | – | 0.449(**) | 0.576 (***) | – | – | 0.374 (*) | – | 0.523 (***) | – | 0.679 (***) | – | – |
| Clay | – | – | –0.775 | 0.449(**) | – | 0.326 (*) | – | – | – | – | 0.727 (***) | – | 0.816 (***) | – | – |
| Texture | – | – | –0.557 | 0.576 (***) | 0.326 (*) | – | – | – | – | – | 0.362 (*) | – | 0.358 (*) | 0.396 (*) | – |
| pH | – | 0.743 (***) | – | – | – | – | – | –0.609 (***) | –0.599 (***) | –0.620 (***) | – | 0.570 (***) | –0.424(**) | 0.602 (***) | 0.506 (***) |
| TOC | – | –0.618 | –0.333 (*) | – | – | – | –0.609 | – | 0.982 (***) | 0.416(**) | – | – | 0.618 (***) | – | – |
| TN | – | –0.637 | –0.396 (*) | 0.374 (*) | – | – | –0.599 | 0.982 (***) | – | 0.316 (*) | – | – | 0.656 (***) | – | – |
| C :N | – | –0.668 | – | – | – | – | –0.620 | 0.416(**) | 0.316 (*) | – | –0.455 | –0.436(**) | – | –0.608 (***) | – |
| SBC | – | – | –0.702 | 0.523 (***) | 0.727 (***) | 0.362 (*) | – | – | – | –0.455(**) | – | 0.531 (***) | 0.720 (***) | 0.661 (***) | – |
| CEC | – | – | – | – | – | 0.396 (*) | 0.602 (***) | – | – | –0.608 | 0.661 (***) | 0.767 (***) | – | – | – |
| BS | – | – | – | – | – | – | 0.506 (***) | – | – | – | – | 0.905 (***) | – | 0.528 (***) | 0.528 (***) |
| Ca :Al | – | 0.338 (*) | – | – | – | – | 0.570 (***) | – | – | –0.436(**) | 0.531 (***) | – | – | 0.767 (***) | – |
| SBC : Al | – | – | –0.853 | 0.679 (***) | 0.816 (***) | 0.358 (*) | –0.424(**) | 0.618 (***) | 0.656 (***) | – | 0.720 (***) | – | – | – | 0.905 (***) |

Figure 4.
Dendrogram of selected physicochemical parameters in different soil types in central and northern Poland (2017)
Table 5.
Valuation of critical acid deposition loads of SBC : Al (A) and Ca : Al (B) of forest soils of centralnorthern Poland in 2007–2017
| A | |||
|---|---|---|---|
| Risk assessment | SBC : Al | Percentage of samples (%) | |
| 2007 | 2017 | ||
| Very High | <0.40 | 50.0 | 50.00 |
| High | 0.40–0.80 | 14.58 | 12.50 |
| Medium | 0.81–2.00 | 8.33 | 4.17 |
| Low | >2.01 | 27.09 | 33.33 |
| B | |||
|---|---|---|---|
| Risk assessment | Ca : Al | Percentage of samples (%) | |
| 2007 | 2017 | ||
| Very high | <0.20 | 45.83 | 47.90 |
| High | 0.21–0.50 | 14.60 | 10.40 |
| Medium | 0.51–1.00 | 6.24 | 10.45 |
| Low | 25.00 | 28.45 | |
1 Soil type (WRB): BRt – Eutric Cambisol, RDb – Brunic Arenosol (Protospodic), BRKt – Dystric Cambisol, OGw – Stagnosol, B – Haplic Podzol, RDbr – Brunic Arenosol, Pbr – Haplic Luvisol, BRg – Gleyic Cambisol; Parent material origin: Qp – glacial sand, QZp – fluvioglacial sand (sandr), Qg – glacial till, QEp/QZp – aeolian sand over fluvioglacial sand (sandr), QFp – fluvioglacial sand; Dominant tree species: So – Scots pine, Bk – European beech, Db – Oak, Św – Norway spruce]; Forest Site Types (TSL): MFCF – mixed fresh coniferous forest; FF – fresh forest; MFF – mixed forest fresh; FCF – fresh coniferous forest