Skip to main content
Have a personal or library account? Click to login
Antimicrobial Resistance in Klebsiella pneumoniae: Mechanisms, Epidemiology, and Emerging Threats Cover

Antimicrobial Resistance in Klebsiella pneumoniae: Mechanisms, Epidemiology, and Emerging Threats

Open Access
|Jul 2026

References

  1. Ambler R.P, Coulson A.F, Frere J.M, Ghuysen J.M, Joris B, Forsman M, Levesque R.C, Tiraby G, Waley S.G. A standard numbering scheme for the class A β–lactamases. Biochemical Journal. 1991;276:269–270. http://doi.org/10.1042/bj2760269
  2. Arakawa Y. Systematic research to overcome newly emerged multidrug–resistant bacteria. Microbiology and Immunology. 2020;64:231–251. http://doi.org/10.1111/1348-0421.12781
  3. Asokan S, Jacob T, Jacob J, AlSosowaa A.A, Cherian T, Peijnenburg W.J.G.M, Vijayan S. Klebsiella pneumoniae: A growing threat in the era of antimicrobial resistance. The Microbe. 2025;7:100333. http://doi.org/10.1016/j.microb.2025.100333
  4. Bader M.S, Loeb M, Leto D, Brooks A.A. Treatment of urinary tract infections in the era of antimicrobial resistance and new antimicrobial agents. Postgraduate Medicine. 2020;32:234–250. http://doi.org/10.1080/00325481.2019.1680052
  5. Bahr G, González L.J, Vila A.J. Metallo-β-lactamases and a tug–of–war for the available zinc at the host-pathogen interface. Current Opinion in Chemical Biology. 2021;66:102103. http://doi.org/10.1016/j.cbpa.2021.102103
  6. Baraniak A, Fiett J, Mrówka A, Walory J, Hryniewicz W, Gniadkowski M. Evolution of TEM-Type Extended–Spectrum β-Lactamases in Clinical Enterobacteriaceae Strains in Poland. Antimicrobial Agents and Chemotherapy. 2005;49:1872–1880. http://doi.org/10.1128/AAC.49.5.1872-1880.2005
  7. Baraniak A & KPC–PL Study Group et al. Molecular characteristics of KPC–producing Enterobacteriaceae at the early stage of their dissemination in Poland, 2008–2009. Antimicrobial Agents and Chemotherapy. 2011;55:5493–5499. http://doi.org/10.1128/AAC.05118-11
  8. Baraniak A & Gniadkowski M et al. NDM–producing Enterobacteriaceae in Poland, 2012–14: inter–regional outbreak of Klebsiella pneumoniae ST11 and sporadic cases. Journal of Antimicrobial Chemotherapy. 2016;71:85–91. http://doi.org/10.1093/jac/dkv282
  9. Baraniak A, Izdebski R, Żabicka D, Bojarska K, Górska S, Literacka E, Fiett J, Hryniewicz W, Gniadkowski M, KPC–PL2 Study Group. Multiregional dissemination of KPC–producing Klebsiella pneumoniae ST258/ST512 genotypes in Poland, 2010–14. Journal of Antimicrobial Chemotherapy. 2017;72:1610–1616. http://doi.org/10.1093/jac/dkx054
  10. Baraniak A, Machulska M, Żabicka D, Literacka E, Izdebski R, Urbanowicz P, Bojarska K, Herda M, Kozinska A, Hryniewicz W, Gniadkowski M, on behalf of the NDM–PL Study Group. Towards endemicity: large–scale expansion of the NDM–1–producing Klebsiella pneumoniae ST11 lineage in Poland, 2015–16. Journal of Antimicrobial Chemotherapy. 2019;74:3199–3204. http://doi.org/10.1093/jac/dkz315
  11. Bauernfeind A, Schneider I, Jungwirth R, Sahly H, Ullmann U. A novel type of AmpC β–lactamase, ACC–1, produced by a Klebsiella pneumoniae strain causing nosocomial pneumonia. Antimicrobial Agents and Chemotherapy. 1999;43:1924–1931. http://doi.org/10.1128/AAC.43.8.1924
  12. Biedrzycka M, Urbanowicz P, Guzek A, Brisse S, Gniadkowski M, Izdebski R. Dissemination of Klebsiella pneumoniae ST147 NDM–1 in Poland, 2015–19. Journal of Antimicrobial Chemotherapy. 2021;76:2538–2545. http://doi.org/10.1093/jac/dkab207
  13. Biedrzycka M, Izdebski R, Urbanowicz P, Polańska M, Hryniewicz W, Gniadkowski M, Literacka E. MDR carbapenemase–producing Klebsiella pneumoniae of the hypervirulence–associated ST23 clone in Poland, 2009–19. Journal of Antimicrobial Chemotherapy. 2022;77:3367–3375. http://doi.org/10.1093/jac/dkac326
  14. Brauncajs M, Bielec F, Macieja A, Pastuszak–Lewandowska D. Carbapenem–resistant Gram–negative fermenting and non–fermenting rods isolated from hospital patients in Poland—what are they susceptible to? Biomedicines. 2022;10:3049. http://doi.org/10.3390/biomedicines10123049
  15. Bose P, Rangnekar A, Desikan P. NDM–β–lactamase–1: where do we stand? Indian Journal of Medical Research. 2022;155:243–252. http://doi.org/10.4103/ijmr.IJMR_685_19
  16. Boyd S.E, Livermore D.M, Hooper D.C, Hope W.W. Metallo–β–lactamases: structure, function, epidemiology, treatment options, and the development pipeline. Antimicrobial Agents and Chemotherapy. 2020;64:e00397. http://doi.org/10.1128/AAC.00397-20
  17. Bradford P.A. Extended–spectrum beta–lactamases in the 21st century: characterization, epidemiology, and detection of this important resistance threat. Clinical Microbiology Reviews. 2001; 14:933–951. http://doi.org/10.1128/CMR.14.4.933-951.2001
  18. Brem J, Schofield Ch.J et al. Imitation of β–lactam binding enables broad–spectrum metallo–β–lactamase inhibitors. Nature Chemistry. 2022 Jan; 14:15–24. http://doi.org/10.1038/s41557-021-00831-x
  19. Bush K, Jacoby G.A. Updated functional classification of β–lactamases. Antimicrobial Agents and Chemotherapy. 2010;54:969–976. http://doi.org/10.1128/AAC.01009-09
  20. Bush K. Past and present perspectives on β–lactamases. Antimicrobial Agents and Chemotherapy. 2018;62:e01076. http://doi.org/10.1128/AAC.01076-18
  21. Bush K, Bradford P.A. Interplay between β–lactamases and new β–lactamase inhibitors. Nature Reviews Microbiology. 2019; 17:295–306. http://doi.org/10.1038/s41579-019-0159-8
  22. Butler D.A, Rana A.P, Krapp F, Patel S.R, Huang Y, Ozer E.A, Hauser A.R, Bulman Z.P. Optimizing aminoglycoside selection for KPC–producing Klebsiella pneumoniae with the aminoglycoside–modifying enzyme (AME) gene aac(6')–Ib. Journal of Antimicrobial Chemotherapy. 2021 Mar; 76:671–679. http://doi.org/10.1093/jac/dkaa480
  23. Calbo E, Garau J. The changing epidemiology of hospital outbreaks due to ESβL–producing Klebsiella pneumoniae: the CTX–M–15 type consolidation. Future Microbiology. 2015;10:1063–1075. https://doi.org/10.2217/fmb.15.22
  24. Caltagirone M, Pagani L et al. Occurrence of extended spectrum β–lactamases, KPC–type, and MCR–1.2–producing Enterobacteriaceae from wells, river water, and wastewater treatment plants in Oltrepò Pavese area, northern Italy. Frontiers in Microbiology. 2017; 8:2232. http://doi.org/10.3389/fmicb.2017.02232
  25. Castanheira M, Simner P.J, Bradford P.A. Extended–spectrum β–lactamases: an update on their characteristics, epidemiology and detection. JAC-Antimicrobial Resistance. 2021 Jul; 3:dlab092. http://doi.org/10.1093/jacamr/dlab092
  26. Catalano A, Iacopetta D, Ceramella J, Scumaci D, Giuzio F, Saturnino C, Aquaro S, Rosano C, Sinicropi M.S. Multidrug resistance (MDR): a widespread phenomenon in pharmacological therapies. Molecules. 2022 Jan; 27:616. http://doi.org/10.3390/molecules27030616
  27. Chambers H.F, Hackbarth C.J. Effect of NaCl and nafcillin on penicillin–binding protein 2a and heterogeneous expression of methicillin resistance in Staphylococcus aureus. Antimicrobial Agents and Chemotherapy. 1987;31:1982–1988. http://doi.org/10.1128/AAC.31.12.1982
  28. Chong Y, Shimoda S, Shimono N. Current epidemiology, genetic evolution and clinical impact of extended–spectrum β–lactamase–producing Escherichia coli and Klebsiella pneumoniae. Infection, Genetics and Evolution. 2018;61:185–188. http://doi.org/10.1016/j.meegid.2018.04.005
  29. Chung P.Y. The emerging problems of Klebsiella pneumoniae infections: carbapenem resistance and biofilm formation. FEMS Microbiology Letters. 2016;363:fnw219. http://doi.org/10.1093/femsle/fnw219
  30. Codjoe F.S, Donkor E.S. Carbapenem resistance: a review. Medical Sciences. 2018;6:1–28. http://doi.org/10.3390/medsci6010001
  31. Council of the European Union. Council Recommendation on stepping up EU actions to combat antimicrobial resistance in a One Health approach (2023/C 220/01). 2023. https://eur-lex.europa.eu/legal-content/EN/TXT/PDF/?uri=OJ:JOC_2023_220_R_0001
  32. Dautzenberg M.J.D, Haverkate M.R, Bonten M.J.M, Bootsma M.C.J. Epidemic potential of Escherichia coli ST131 and Klebsiella pneumoniae ST258: a systematic review and meta–analysis. BMJ Open. 2016;6:e009971. http://doi.org/10.1136/bmjopen-2015-009971
  33. Ding L, Shen S, Chen J, Tian Z, Shi Q, Han R, Guo Y, Hu F. Klebsiella pneumoniae carbapenemase variants: the new threat to global public health. Clinical Microbiology Reviews. 2023;36:e00008-23. http://doi.org/10.1128/cmr.00008-23
  34. Dzierżanowska D. Antybiotykoterapia praktyczna. Alfa-Medica Press; 6th edition. 2024;1–752.
  35. Dzierżanowska-Fangrat. Antibiotic Therapy Guide 2023. Alfa Medica Press. 2023;1–132.
  36. Dzierżanowska D, Kamińska W, Semczuka K, Borowiec D, Matysiak M, Szumała–Kąkol A, Gierczyński R, Patzer J.A. Carriage of genes for various extended–spectrum β–lactamases: a novel resistance strategy of Klebsiella pneumoniae in Poland. International Journal of Antimicrobial Agents. 2010;35:392–395. http://doi.org/10.1016/j.ijantimicag.2009.12.010
  37. Dziewit Ł, Bartosik D. Prokaryotic genomes in the light of genomic analyses. Advances in Microbiology. 2011;50:87–96.
  38. European Centre for Disease Prevention and Control (ECDC). ECDC Surveillance Atlas of Infectious Diseases. https://www.ecdc.europa.eu
  39. Esumeh F.I, Inyang N.J, Adesina I.A, Akpe A.R, Omoigberale M.N.O, Igunbor L.A. Antibacterial susceptibility and in vitro effect of sodium chloride, magnesium ions, and serum on staphylococcal β–lactamase production in Ekpoma, Nigeria. Special Bacteriology Pathogen Journal. 2017;2:01–07. http://doi.org/10.61915/bpj.600192
  40. European Centre for Disease Prevention and Control. European Antimicrobial Resistance Surveillance Network (EARS–Net). https://ecdc.europa.eu/en/about-us/networks/disease-networks-and-laboratory-networks/ears-netabout
  41. European Centre for Disease Prevention and Control. Antimicrobial resistance in the EU/EEA (EARS–Net) – Annual Epidemiological Report 2024. Stockholm: ECDC. https://www.ecdc.europa.eu/en/publications-data/antimicrobial-resistance-eueea-ears-net-annual-epidemiological-report-2024
  42. European Centre for Disease Prevention and Control. Antimicrobial resistance in the EU/EEA (EARS–Net) – Surveillance of antimicrobial resistance in Europe, 2024 data. Stockholm: ECDC. https://www.ecdc.europa.eu/sites/default/files/documents/antimicrobial-resistance-surveillance-Europe-2024-data.pdf
  43. Fevre C, Passet V, Weill F.-X, Patrick A, Grimont D, Brisse S. Variants of the Klebsiella pneumoniae OKP chromosomal β–lactamase are divided into two main groups, OKP–A and OKP–B. Journal of Clinical Microbiology. 2005;43:4178–4182. http://doi.org/10.1128/JCM.43.8.4178-4182.2005
  44. Fiett J, Baraniak A, Izdebski R, Sitkiewicz I, Żabicka D, Meler A, Filczak K, Hryniewicz W, Gniadkowski M. The first NDM metallo–β–lactamase–producing Enterobacteriaceae isolate in Poland: evolution of IncFII–type plasmids carrying the blaNDM–1 gene. Antimicrobial Agents and Chemotherapy. 2014;58:1203–1207. http://doi.org/10.1128/aac.01197-13
  45. Galani I, Karaiskos I, Giamarellou H. Multidrug–resistant Klebsiella pneumoniae: mechanisms of resistance including updated data for novel β–lactam–β–lactamase inhibitor combinations. Expert Review of Anti-Infective Therapy. 2022;19:1457–1468. http://doi.org/10.1080/14787210.2021.1924674
  46. Gniadkowski M, Palucha A, Grzesiowski P, Hryniewicz W. Outbreak of ceftazidime–resistant Klebsiella pneumoniae in a pediatric hospital in Warsaw, Poland: clonal spread of the TEM–47 extended–spectrum β–lactamase (ESβL)–producing strain and transfer of a plasmid carrying the SHV–5–like ESβL–encoding gene. Antimicrobial Agents and Chemotherapy. 1998;42:3079–3085. http://doi.org/10.1128/AAC.42.12.3079
  47. Gniadkowski M, Schneider I, Jungwirth R, Hryniewicz W, Bauernfeind A. Ceftazidime–resistant Enterobacteriaceae isolates from three Polish hospitals: identification of three novel TEM– and SHV–5–type extended–spectrum β–lactamases. Antimicrobial Agents and Chemotherapy. 1998;42:514–520. http://doi.org/10.1128/aac.42.3.514
  48. Guo L, An J, Ma Y, Ye L, Luo Y, Tao C, Yang J. Nosocomial outbreak of OXA–48–producing Klebsiella pneumoniae in a Chinese hospital: clonal transmission of ST147 and ST383. PLOS ONE. 2016;11:e0160754. http://doi.org/10.1371/journal.pone.0160754
  49. Guzek A, Rybicki Z, Tomaszewski D, Mackiewicz K, Piechota W, Chciałowski A. Outcomes of 23 patients diagnosed with New Delhi metallo–β–lactamase (NDM)–producing Klebsiella pneumoniae infection treated with ceftazidime/avibactam and aztreonam at a single center in Poland. European Journal of Clinical Microbiology & Infectious Diseases. 2024;43:1579–1587. http://doi.org/10.1007/s10096-024-04859-y
  50. Hæggman S, Löfdahl S, Paauw A, Verhoef J, Brisse S. Diversity and evolution of the class A chromosomal β–lactamase gene in Klebsiella pneumoniae. Antimicrobial Agents and Chemotherapy. 2004;48:2400–2408. http://doi.org/10.1128/AAC.48.7.2400-2408.2004
  51. Huang L, Wu C, Gao H, Xu C, Dai M, Huang L, Hao H, Wang X, Cheng G. Bacterial multidrug efflux pumps at the frontline of antimicrobial resistance: an overview. Antibiotics (Basel). 2022;11:520. http://doi.org/10.3390/antibiotics11040520
  52. Hughes S, Gilchrist M, Heard K, Hamilton R, Sneddon J. Treating infections caused by carbapenemase–producing Enterobacterales (CPE): a pragmatic approach to antimicrobial stewardship on behalf of the UKCPA Pharmacy Infection Network (PIN). JAC-Antimicrobial Resistance. 2020;2:dlaa075. http://doi.org/10.1093/jacamr/dlaa075
  53. Ishizaki Y, Shibuya Y, Hayashi C, Inoue K, Kirikae T, Tada T, Miyoshi–Akiyama T, Igarashi M. Instability of the 16S rRNA methyltransferase–encoding npmA gene: why have bacterial cells possessing npmA not spread despite their high and broad resistance to aminoglycosides? Journal of Antibiotics. 2018;1:798–807. http://doi.org/10.1038/s41429-018-0070-y
  54. Izdebski R & OXA–48–PL Study Group et al. Enterobacteriaceae producing OXA–48–like carbapenemases in Poland, 2013–January 2017. Journal of Antimicrobial Chemotherapy. 2018;73:620–625. http://doi.org/10.1093/jac/dkx457
  55. Jacoby G.A. AmpC β–lactamases. Clinical Microbiology Reviews. 2009;22:161–182. http://doi.org/10.1128/CMR.00036-08
  56. Journal of Laws of the Republic of Poland. Regulation of the Council of Ministers of March 30, 2021, on the National Health Program for 2021–2025. Warsaw, April 8, 2021, Item 642. KORLD.
  57. Karami–Zarandi M, Rahdar H.A, Esmaeili H, Ranjbar R. Klebsiella pneumoniae: an update on antibiotic resistance mechanisms. Future Microbiology. 2023;18:65–81. http://doi.org/10.2217/fmb-2022-0097
  58. Khattab S, Askar A.M, Abdellatif H.A.A, Othman A.A.A, Rayan A.H, Azab H. Synergistic combination of ceftazidime and avibactam with aztreonam against MDR Klebsiella pneumoniae in ICU patients. Scientific Reports. 2025;15:5102. http://doi.org/10.1038/s41598-025-88965-7
  59. Kiener P.A, Knott–Hunziker V, Petursson S, Waley S.G. Mechanism of substrate–induced inactivation of β–lactamase I. European Journal of Biochemistry. 1980;109:575–580. http://doi.org/10.1111/j.1432-1033.1980.tb04830.x
  60. Koch A.L. Autolysis control hypotheses for tolerance to wall antibiotics. Antimicrobial Agents and Chemotherapy. 2001;45:2671–2675. http://doi.org/10.1128/AAC.45.10.2671-2675.2001
  61. Kowalska–Krochmal B, Woroń J, Serednicki W.T, Wordliczek J. Ceftazidime–avibactam – microbiological, pharmacological and clinical aspects in clinical practice. Forum Infectiology. 2019;10:373–381. http://doi.org/10.15374/FZ2019056
  62. Krajewska J, Laudy A.E. The European Medicines Agency approved the new antibacterial drugs – response to the 2017 WHO report on the global problem of multi–drug resistance. Advances in Microbiology. 2021;60:249–264. http://doi.org/10.21307/pm-2021.60.4.20
  63. Kramer A, Schwebke I, Kampf G. How long do nosocomial pathogens persist on inanimate surfaces? A systematic review. BMC Infectious Diseases. 2006;6:130. http://doi.org/10.1186/1471-2334-6-130
  64. Krul D, Dalla Costa M.L. High–risk clones of carbapenem–resistant Klebsiella pneumoniae recovered from pediatric patients in Southern Brazil. Brazilian Journal of Microbiology. 2024;55:1437–1443. http://doi.org/10.1007/s42770-024-01299-w
  65. Kuch A, Hryniewicz W, Wanke–Rytt M, Żukowska A. Pałeczki Enterobacterales wytwarzające karbapenemazy (CPE): epidemiologia, diagnostyka, leczenie i profilaktyka zakażeń. Warszawa: Narodowy Instytut Leków; 2022.
  66. Laishram S, Anandan S, Devi B.Y, Elakkiya M, Priyanka B, Bhuvaneshwari T, Peter J.V, Subramani K, Balaji V. Determination of synergy between sulbactam, meropenem and colistin in carbapenem–resistant Klebsiella pneumoniae and Acinetobacter baumannii isolates and correlation with the molecular mechanism of resistance. Journal of Chemotherapy. 2016;28:297–303. http://doi.org/10.1179/1973947815Y.0000000079
  67. Lee C.R, Lee J.H, Park K.S, Kim Y.B, Jeong B.Ch, Lee S.H. Global dissemination of carbapenemase–producing Klebsiella pneumoniae: epidemiology, genetic context, treatment options, and detection methods. Frontiers in Microbiology. 2016;7:1–30. http://doi.org/10.3389/fmicb.2016.00895
  68. Lei T.–Y, Liao B.–B, Yang L.–R, Wang Y, Chen X.–B. Hypervirulent and carbapenem–resistant Klebsiella pneumoniae: a global public health threat. Microbiological Research. 2024;288:127839. http://doi.org/10.1016/j.micres.2024.127839
  69. Li Y, Kumar S, Zhang L. Mechanisms of antibiotic resistance and developments in therapeutic strategies to combat Klebsiella pneumoniae infection. Infection and Drug Resistance. 2024;17:1107–1119. http://doi.org/10.2147/IDR.S453025
  70. Liakopoulos A, Dik Mevius D, Ceccarelli D. A review of SHV extended–spectrum β–lactamases: neglected yet ubiquitous. Frontiers in Microbiology. 2016;7:1374. http://doi.org/10.3389/fmicb.2016.01374
  71. Litwin A, Fedorowicz O, Duszyńska W. Characteristics of microbial factors of healthcare–associated infections including multidrug–resistant pathogens and antibiotic consumption at the University Intensive Care Unit in Poland in the years 2011–2018. International Journal of Environmental Research and Public Health. 2020;17:6943. http://doi.org/10.3390/ijerph17196943
  72. Liu C, Wu Y, Fang Y, Sang Z, Huang L, Dong N, Zeng Y, Lu J, Zhang R, Chen G. Emergence of an ST1326 (CG258) multi-drug–resistant Klebsiella pneumoniae co–harboring mcr–8.2, ESβL genes, and the resistance–nodulation–division efflux pump gene cluster tmexCD1–toprJ1 in China. Frontiers in Microbiology. 2022;13:800993. http://doi.org/10.3389/fmicb.2022.800993
  73. Livermore D.M. β–lactamases in laboratory and clinical resistance. Clinical Microbiology Reviews. 1995;8:557–584. http://doi.org/10.1128/CMR.8.4.557
  74. Loconsole D, Accogli M, De Robertis A.L, Capozzi L, Bianco A, Morea A, Mallamaci R, Quarto M, Parisi A, Chironna M. Emerging high–risk ST101 and ST307 carbapenem–resistant Klebsiella pneumoniae clones from bloodstream infections in Southern Italy. Annals of Clinical Microbiology and Antimicrobials. 2020;19:24. http://doi.org/10.1186/s12941-020-00366-y
  75. Machulska M, Baraniak A, Żak I, Bojarska K, Żabicka D, Sowa–Sierant I, Hryniewicz W, Gniadkowski M. KPC–2–producing Klebsiella pneumoniae ST11 in a children's hospital in Poland. Polish Journal of Microbiology. 2017;66:401–404. http://doi.org/10.5604/01.3001.0010.4884
  76. Mączyńska B, Neumann K, Junka A. Analysis of properties related to selection and survival in hospital environment of Klebsiella strains isolated from nosocomial outbreaks. Forum Infectiology. 2015;4:77–97. http://doi.org/10.15374/fz2013013
  77. Mączyńska B. Evolution of pathogenicity and resistance to antibacterial agents in Klebsiella bacilli. Evereth Publishing Sp. z o.o. 2015.
  78. McCreary E.K, Heil E.L, Tamma P.D. New perspectives on antimicrobial agents: cefiderocol. Antimicrobial Agents and Chemotherapy. 2021;65:e02171-20. http://doi.org/10.1128/AAC.02171-20
  79. Mehta S.C, Rice K, Patel T. Natural variants of the KPC–2 carbapenemase have evolved increased catalytic efficiency for ceftazidime hydrolysis at the cost of enzyme stability. PLoS Pathogens. 2015;11:e1004949. http://doi.org/10.1371/journal.ppat.1004949
  80. Meini S, Tascini C, Cei M, Sozio E, Rossolini G.M. AmpC β–lactamase–producing Enterobacterales: what a clinician should know. Infection. 2019;47:363–375. http://doi.org/10.1007/s15010-019-01291-9
  81. Meek R.W, Vyas H, Piddock L.J.V. Nonmedical uses of antibiotics: time to restrict their use? PLoS Biology. 2015;13:1–11. http://doi.org/10.1371/journal.pbio.1002266
  82. Musawa M.A, Bleick C.R, Herbin S.R, Caniff K.E, Van Helden S.R, Rybak M.J. Aztreonam–avibactam: the dynamic duo against multidrug–resistant Gram–negative pathogens. Pharmacotherapy. 2024;44:927–938. http://doi.org/10.1002/phar.4629
  83. Müderris T, Manyaslı G.D, Kaya S, Yurtsever S.G. In vitro interactions of combinations of colistin with meropenem, rifampicin and tigecycline in colistin–resistant, biofilm–forming Klebsiella pneumoniae. Diagnostic Microbiology and Infectious Disease. 2024;110:116408. http://doi.org/10.1016/j.diagmicrobio.2024.116408
  84. Navon–Venezia S, Kondratyeva K, Carattoli A. Klebsiella pneumoniae: a major worldwide source and shuttle for antibiotic resistance. FEMS Microbiology Reviews. 2017;41:252–275. http://doi.org/10.1093/femsre/fux013
  85. Nordmann P, Poirel L. The difficult–to–control spread of carbapenemase producers among Enterobacteriaceae worldwide. Clinical Microbiology and Infection. 2014;20:821–830. http://doi.org/10.1111/1469-0691.12719
  86. Ochońska D, Brzychczy–Włoch M. Klebsiella pneumoniae – taxonomy, occurrence, identification, virulence factors and pathogenicity. Advances in Microbiology. 2024;63:157–175. http://doi.org/10.2478/am-2024-0014
  87. Ochońska D, Klamińska–Cebula H, Dobrut A, Bulanda M, Brzychczy–Włoch M. Clonal dissemination of KPC–2, VIM–1, OXA–48–producing Klebsiella pneumoniae ST147 in Katowice, Poland. Polish Journal of Microbiology. 2021;70:107–116. http://doi.org/10.33073/pjm-2021-010
  88. Ojdana D, Sacha P, Olszańska D, Majewski P, Wieczorek P, Jaworowska J, Sieńko A, Jurczak A, Tryniszewska E. First report of Klebsiella pneumoniae carbapenemase–3–producing Escherichia coli ST479 in Poland. BioMed Research International. 2015;2015:256028. http://doi.org/10.1155/2015/256028
  89. Papanicolaou G.A, Medeiros A.A, Jacoby G.A. Novel plasmid–mediated β–lactamase (MIR–1) conferring resistance to oxyimino– and o–methoxy β–lactams in clinical isolates of Klebsiella pneumoniae. Antimicrobial Agents and Chemotherapy. 1990;34:2200–2209. http://doi.org/10.1128/aac.34.11.2200
  90. Parra Sellera F, Lincopan N, Fuentes–Castillo D, Guedes Stehling E, Rueda Furlan J.P. Rapid evolution of pan–β–lactam resistance in Enterobacterales co–producing KPC and NDM: insights from global genomic analysis after the COVID–19 pandemic. The Lancet Microbe. 2024;5:e412–e413. http://doi.org/10.1016/S2666-5247(24)00018-1
  91. Paterson D.L, Hujer K.M, Hujer A.M, Yeiser B, Bonomo M.D, Rice L.B, Bonomo R.A, International Klebsiella Study Group. Extended–spectrum β–lactamases in Klebsiella pneumoniae bloodstream isolates from seven countries: dominance and widespread prevalence of SHV– and CTX–M–type β–lactamases. Antimicrobial Agents and Chemotherapy. 2003;47:3554–3560. http://doi.org/10.1128/AAC.47.11.3554-3560.2003
  92. Paul M, Rodríguez–Baño J. et al. European Society of Clinical Microbiology and Infectious Diseases (ESCMID) guidelines for the treatment of infections caused by multidrug–resistant Gram–negative bacilli (endorsed by European Society of Intensive Care Medicine). Clinical Microbiology and Infection. 2022;28:521–547. http://doi.org/10.1016/j.cmi.2021.11.025
  93. Peirano G, Chen L, Kreiswirth B.N, Pitout J.D.D. Emerging antimicrobial–resistant high–risk Klebsiella pneumoniae clones ST307 and ST147. Antimicrobial Agents and Chemotherapy. 2020;64:e01148-20. http://doi.org/10.1128/AAC.01148-20
  94. Philippon A, Redjeb S.B, Fournier G, Hassen A.B. Epidemiology of extended–spectrum β–lactamases. Infection. 1989;17:347–354. http://doi.org/10.1007/BF01650727
  95. Pitout J.D.D, Nordmann P, Poirel L. Carbapenemase–producing Klebsiella pneumoniae, a key pathogen set for global nosocomial dominance. Antimicrobial Agents and Chemotherapy. 2015;59:5873–5884. http://doi.org/10.1128/AAC.01019-15
  96. Pitout J.D.D, Peirano G, Kock M.M, Strydom K.–A, Matsumura Y. The global ascendency of OXA–48–type carbapenemases. Clinical Microbiology Reviews. 2019;33:e00102-19. http://doi.org/10.1128/CMR.00102-19
  97. Pitton J.S. Mechanism of bacterial resistance to antibiotics. Reviews of Physiology. 1972;65:15–93. http://doi.org/10.1007/3-540-05814-1_2
  98. Polish National Reference Centre for Antimicrobial Susceptibility Testing (KORLD). https://korld.nil.gov.pl/, https://korld.nil.gov.pl/opornosc-na-antybiotyki-2/raporty-korld/
  99. Rajan R, Sasikala G. Evaluation of the in vitro efficacy of anti-microbials against Enterobacterales with multiple carbapenemase enzymes. Iranian Journal of Microbiology. 2025;17:390–396. http://doi.org/10.18502/ijm.v17i3.18821
  100. State Sanitary Inspection (Poland). Report on the Activities of the State Sanitary Inspection in the Field of Public Health for 2024. https://www.gov.pl/web/gis/raport-stan-sanitarny-kraju
  101. Rocker A, Lacey J.A, Belousoff M.J, Wilksch J.J, Strugnell R.A, Davies M.R, Lithgow T. Global trends in proteome remodeling of the outer membrane modulate antimicrobial permeability in Klebsiella pneumoniae. mBio. 2020;11:e00603-20. http://doi.org/10.1128/mBio.00603-20
  102. Rodríguez–Guerrero E, Callejas–Rodelas J.C, Navarro–Marí J.M, Gutiérrez–Fernández J. Systematic review of plasmid AmpC type resistances in Escherichia coli and Klebsiella pneumoniae and preliminary proposal of a simplified screening method for ampC. Microorganisms. 2022;10:611. http://doi.org/10.3390/microorganisms10030611
  103. Sarowska J, Choroszy–Krol I, Jama–Kmiecik A, Mączyńska B, Cholewa S, Frej–Madrzak M. Occurrence and characteristics of carbapenem–resistant Klebsiella pneumoniae strains isolated from hospitalized patients in Poland – a single centre study. Pathogens. 2022;11:859. http://doi.org/10.3390/pathogens11080859
  104. Sawa T, Kooguchi K, Moriyama K. Molecular diversity of extended–spectrum β–lactamases and carbapenemases, and antimicrobial resistance. Journal of Intensive Care. 2020;8:13. http://doi.org/10.1186/s40560-020-0429-6
  105. Sękowska A. In vitro activity of “old” and “new” antimicrobials against the Klebsiella pneumoniae complex. Antibiotics (Basel). 2024;13:126. http://doi.org/10.3390/antibiotics13020126
  106. Sękowska A. In vitro activity of plazomicin and other aminoglycosides against Klebsiella pneumoniae multidrug–resistant strains. Journal of Antibiotics. 2024;77:548–551. http://doi.org/10.1038/s41429-024-00734-2
  107. Shah A.A, Alwashmi A.S.S, Abalkhail A, Alkahtani A.M. Emerging challenges in Klebsiella pneumoniae: antimicrobial resistance and novel approaches. Microbial Pathogenesis. 2025;202:107399. http://doi.org/10.1016/j.micpath.2025.107399
  108. Shortridge D, Streit J.M, Mendes R, Castanheira M. In vitro activity of cefiderocol against U.S. and European Gram–negative clinical isolates collected in 2020 as part of the SENTRY antimicrobial surveillance program. Microbiology Spectrum. 2021;10:e02712-21. http://doi.org/10.1128/spectrum.02712-21
  109. Sinha R, Khare S.K. Protective role of salt in catalysis and maintaining structure of halophilic proteins against denaturation. Frontiers in Microbiology. 2014;5:165. http://doi.org/10.3389/fmicb.2014.00165
  110. Slain N, Lucas K, Kunz Coyne A.J. Comparison between cefepime and carbapenem therapy for deep–seated AmpC–producing Enterobacterales infections: a propensity–weighted retrospective cohort study. Antimicrobial Agents and Chemotherapy. 2025;69:e00928-25. http://doi.org/10.1128/aac.00928-25
  111. Słabisz N, Leśnik P, Janc J, Fidut M, Bartoszewicz M, Dudek–Wicher R, Nawrot U. Evaluation of the in vitro susceptibility of clinical isolates of NDM–producing Klebsiella pneumoniae to new antibiotics included in a treatment regimen for infections. Frontiers in Microbiology. 2024;15:1331628. http://doi.org/10.3389/fmicb.2024.1331628
  112. Stewart A, Harris P, Henderson A, Paterson D. Treatment of infections by OXA–48–producing Enterobacteriaceae. Antimicrobial Agents and Chemotherapy. 2018;62:e01195-18. http://doi.org/10.1128/AAC.01195-18
  113. Subramanian G.K, Soundari P.G, Ramanathan V, Krishnan P. Endemic Indian clones of Klebsiella pneumoniae harbouring New Delhi metallo–β–lactamase–1 on a hybrid plasmid replicon type: a case of changing NDM plasmid landscapes in India? Indian Journal of Medical Microbiology. 2016;34:286–292. http://doi.org/10.4103/0255-0857.188314
  114. Syed Y.Y. Cefiderocol: a review in serious Gram–negative bacterial infections. Drugs. 2021;81:1559–1571. http://doi.org/10.1007/s40265-021-01580-4
  115. Szewczyk E.M. Bacteriological diagnostics. Warsaw: Państwowe Wydawnictwo Naukowe (PWN). 2019;3:1–500.
  116. Szymański M, Skiba M.M, Piasecka M, Olender A. Synergistic effect of ceftazidime–avibactam with aztreonam on carbapenemase–positive Klebsiella pneumoniae MBL+, NDM. Infection and Drug Resistance. 2024;17:2307–2313. http://doi.org/10.2147/IDR. S459695
  117. Ślusarz K, Kurdyś P, Łanowy P, Bichalski M, Bijak B, Trejnowska E. Venovenous extracorporeal membrane oxygenation (VV ECMO) in community–acquired pneumonia caused by Klebsiella pneumoniae: what went wrong? Journal of Education, Health and Sport. 2019;9:738–746. http://doi.org/10.5281/zenodo.3407541
  118. Tamma P.D, Doi Y, Bonomo R.A, Johnson J.K, Simner P.J, Antibacterial Resistance Leadership Group. A primer on AmpC β–lactamases: necessary knowledge for an increasingly multidrug–resistant world. Clinical Infectious Diseases. 2019;69:1446–1455. http://doi.org/10.1093/cid/ciz173
  119. Tarski I, Śmiechowicz J, Duszyńska W. Cefiderocol in the successful treatment of complicated hospital–acquired Klebsiella pneumoniae NDM, OXA–48 intraabdominal infection. Infection and Drug Resistance. 2024;17:5163–5170. http://doi.org/10.2147/IDR. S485450
  120. Tebano G, Zaghi I, Cricca M, Cristini F. Antibiotic treatment of infections caused by AmpC–producing Enterobacterales. Pharmacy (Basel). 2024;12:142. http://doi.org/10.3390/pharmacy12050142
  121. Tsang K.K, KlebNETGSP AMR Genotype–Phenotype Group et al. Diversity, functional classification and genotyping of SHV β–lactamases in Klebsiella pneumoniae. Microbial Genomics. 2024;10:001294. http://doi.org/10.1099/mgen.0.001294
  122. Wang Y, Zhang Q, Jin Y, Jin X, Yu J, Wang K. Epidemiology and antimicrobial susceptibility profiles of extended–spectrum β–lactamase–producing Klebsiella pneumoniae and Escherichia coli in China. Brazilian Journal of Microbiology. 2019;50:669–675. http://doi.org/10.1007/s42770-019-00081-7
  123. Wantuch P.L, Rosen D.A. Klebsiella pneumoniae: adaptive immune landscapes and vaccine horizons. Trends in Immunology. 2023;44:826–844. http://doi.org/10.1016/j.it.2023.08.005
  124. World Health Organization (WHO). Antimicrobial resistance surveillance in Europe 2023 – 2021 data. Copenhagen: WHO Regional Office for Europe; 2025. https://www.who.int/
  125. World Health Organization (WHO). Global Antimicrobial Resistance and Use Surveillance System (GLASS). 2025. https://www.who.int/initiatives/glass
  126. Wyres K.L, Holt K.E. Klebsiella pneumoniae as a key trafficker of drug resistance genes from environmental to clinically important bacteria. Current Opinion in Microbiology. 2018;45:131–139. http://doi.org/10.1016/j.mib.2018.04.004
  127. Wyres K.L, Lam M.M.C, Holt K.E. Population genomics of Klebsiella pneumoniae. Nature Reviews Microbiology. 2020;18:344–359. http://doi.org/10.1016/j.mib.2018.04.004
  128. Wysocka M, Zamudio R, Oggioni M.R, Gołębiewska J, Bronk M, Krawczyk B. Genetic background and antibiotic resistance profiles of Klebsiella pneumoniae NDM–1 strains isolated from UTI, ABU, and the GI tract from one hospital in Poland in relation to strains nationally and worldwide. Genes. 2021;12:1285. http://doi.org/10.3390/genes12081285
  129. Yahav D, Giske C.G, Grāmatniece A, Abodakpi H, Tam V.H, Leibovici L. New β–lactam–β–lactamase inhibitor combinations. Clinical Microbiology Reviews. 2020;34:e00115-20. http://doi.org/10.1128/CMR.00115-20
  130. Yang X, Sun Q, Li J, Jiang Y, Li Y, Lin J, Chen K, Chan E.W.C, Zhang R, Chen S. Molecular epidemiology of carbapenem–resistant hypervirulent Klebsiella pneumoniae in China. Emerging Microbes & Infections. 2022;11:841–849. http://doi.org/10.1080/22221751.2022.2049458
  131. van Duin D, Cober E, Richter S.S, Perez F, Kalayjian R.C, Salata R.A, Evans S, Fowler V.G Jr, Bonomo R.A, Kaye K.S. Residence in skilled nursing facilities is associated with tigecycline non–susceptibility in carbapenem–resistant Klebsiella pneumoniae. Infection Control & Hospital Epidemiology. 2015;36:942–948. http://doi.org/10.1017/ice.2015.118
  132. van Duin D, Paterson D.L. Multidrug–resistant bacteria in the community: an update. Infectious Disease Clinics of North America. 2020;34:709–722. http://doi.org/10.1016/j.idc.2020.08.002
  133. Yoon E–J, Jeong S.H. Class D β–lactamases. Journal of Antimicrobial Chemotherapy. 2021;76:836–864. http://doi.org/10.1093/jac/dkaa513
  134. Yuan P.–B, Dai L.–T, Zhang Q.–K, Zhong Y.–X, Liu W.–T, Yang L, Chen D.–Q. Global emergence of double and multi–carbapenemase producing organisms: epidemiology, clinical significance, and evolutionary benefits on antimicrobial resistance and virulence. Microbiology Spectrum. 2024;12:e00008-24. http://doi.org/10.1128/spectrum.00008-24
DOI: https://doi.org/10.2478/am-2026-0006 | Journal eISSN: 2545-3149 | Journal ISSN: 0079-4252
Language: English, Polish
Page range: 56 - 80
Submitted on: Apr 30, 2025
Accepted on: May 19, 2026
Published on: Jul 28, 2026
In partnership with: Paradigm Publishing Services
Publication frequency: 4 issues per year

© 2026 Dorota Ochońska, Monika Brzychczy–Włoch, published by Polish Society of Microbiologists
This work is licensed under the Creative Commons Attribution-NonCommercial-NoDerivatives 4.0 License.