References
- 1. Friendship, R. M., Melnichouk, S. I., Dewey, C. E., 2000: Omeprazole to alleviate stomach ulcers during feed withdrawal. Can. Vet. J., 41, 925–928.
- 2. Gottardo, F., Scollo, A., Contiero, B., Bottacini, M., Mazzoni, C., Edwards, S. A., 2017: Prevalence and risk factors for gastric ulceration in pigs slaughtered at 170 kg. Animal, 11, 11, 2030–2038. DOI: 10.1017/S1751731117000799
- 3. Pluske, J. R., Turpin, D. L., Kim, J. C., 2018: Gastrointestinal tract (gut) health in the young pig. Anim. Nutr., 4, 2, 187–196. DOI: 10.1016/j.aninu.2017.12.004
- 4. Cybulski, P., Wo ź niak, A., Larska, M., Jab ł o ń ski, A., Stadejek, T., 2024: Gastric ulcers in finishing pigs: the evaluation of selected nondietary risk factors and impact on production performance. Porcine Health Manag., 10, 11. DOI: 10.1186/s40813-024-00362-0
- 5. Roels, S., Ducatelle, R., Broekaert, D., 1997: Keratin pattern in hyperkeratotic and ulcerated gastric pars oesophagea in pigs. Res. Vet. Sci., 62, 2, 165–169. DOI: 10.1016/S0034-5288(97)90140-5
- 6. Pej č inovska, N., Dodovski, P., Karabolovski, N., Zdraveski, I., Dameski, P., Hristovska, T., et al. 2020: Morphological and histological features of the stomach in domestic pigs (Sus scrofa domestica). J. Hyg. Eng. Des., 50, 50–55. https://keypublishing.org/jhed/wp-content/uploads/2025/06/04.-Full-paper-Natasa-Pejcinovska-1.pdf
- 7. Ballari, S. A., Barrios-Garc í a, M. N., 2014: A review of wild boar Sus scrofa diet and factors affecting food selection in native and introduced ranges. Mammal Rev., 44, 2, 124–134. DOI: 10.1111/mam.12015
- 8. Montes-S á nchez, J. J., Huato-Soberanis, L., Buntinx-Dios, S. E., Le ó n-de la Luz, J. L., 2020: The feral pig in a low impacted ecosystem: analysis of diet composition and its utility. Rangel. Ecol. Manag., 73, 5, 703–711. DOI: 10.1016/j.rama.2020.05.006
- 9. Racewicz, P., Ludwiczak, A., Skrzypczak, E., Sk ł adanowskaBaryza, J., Biesiada, H., Nowak, T., et al., 2021: Welfare Health and Productivity in Commercial Pig Herds. Animals, 11, 4, 1176. DOI: 10.3390/ani11041176
- 10. Pinna, W., Nieddu, G., Moniello, G., Cappai, M. G., 2007: Vegetable and animal food sorts found in the gastric content of Sardinian wild boar (Sus scrofa meridionalis). J. Anim. Physiol. Anim. Nutr., 91, 252–255. DOI: 10.1111/j.1439-0396.2007.00700.x
- 11. Zanoni, R.G., Piva, S., Florio, D., Bassi, P., Mion, D., Cnockaert, M., et al. 2016. Helicobacter apri sp. nov., isolated from wild boars. Int. J. Syst. Evol. Microbiol., 66, 8, 2876–2882. DOI:10.1099/ijsem.0.001071
- 12. Cortez Nunes, F., Letra Mateus, T., Teixeira, S., Barradas, P., de Witte, C., Haesebrouck, F., et al., 2021: Presence of Helicobacter pylori and H. suis DNA in free-range wild boars. Animals, 11, 5, 1269. DOI: 10.3390/ani11051269
- 13. Uhr, G., 1995: The intestinal tract and the Peyer’s patch dimensions of wild boars (Sus scrofa L., 1758) and domestic pigs (Sus scrofa f. domestica). An allometric comparison. J. Mt. Ecol., 3, 77–82.
- 14. Wondra, K. J., Hancock, J. D., Behnke, K. C., Hines, R. H., Stark, C. R., 1995: Effects of particle size and pelleting on growth performance, nutrient digestibility, and stomach morphology in finishing pigs. J. Anim. Sci., 73, 3, 757–763. DOI: 10.2527/1995.733757x
- 15. Su, Y., Yao, W., Perez-Gutierrez, O. N., Smidt, H., Zhu, W. Y. 2008: Changes in abundance of Lactobacillus spp. and Streptococcus suis in the stomach, jejunum and ileum of pig-lets after weaning. FEMS Microbiol. Ecol., 66, 3, 546–555. DOI: 10.1111/j.1574-6941.2008.00529.x
- 16. Cappai, M. G., Picciau, M., Pinna, W., 2013: Ulcerogenic risk assessment of diets for pigs in relation to gastric lesion prevalence. BMC Vet. Res., 9, 36, 1–8. DOI: 10.1186/1746-6148-9-36
- 17. Liermann, W., Berk, A., B ö schen, V., D ä nicke, S., 2015: Effects of particle size and hydro-thermal treatment of feed on performance and stomach health in fattening pigs. Arch. Anim. Nutr., 69, 6, 455–472. DOI: 10.1080/1745039X.2015.1087748
- 18. Peralvo-Vidal, J. M., Weber, N. R., Nielsen, J. P., Bache, J. K., Haugegaard, S., Pedersen, A. O., 2021: Risk factors for gastric ulceration in nursery pigs. Prev. Vet. Med., 189, 105298. DOI: 10.1016/j.prevetmed.2021.105298
- 19. Herrero, J., Irizar, I., Laskurain, N.A., Garc í a-Serrano, A., Garc í a-Gonz á lez, R. 2005: Fruits and roots: wild boar foods during the cold season in the southwestern Pyrenees. Ital. J. Zool., 72, 1, 49–52. DOI: 10.1080/11250000509356652
- 20. Taillieu, E., Chiers, K., Amorim, I., G ä rtner, F., Maes, D., Van Steenkiste, C., et al., 2022: Gastric Helicobacter species associated with dogs, cats and pigs: significance for public and animal health. Vet. Res., 53, 42, 1–15. DOI: 10.1186/s13567-022-01059-4
- 21. Camerlink, I., Turner, S. P., Ursinus, W. W., Reimert, I., Bolhuis, J. E., 2014: Aggression and affiliation during social conflict in pigs. PLOS One, 9, 11, e113502. DOI: 10.1371/journal.pone.0113502
- 22. Herskin, M. S., Jensen, H. E., Jespersen, A., Forkman, B., Jensen, M.B., Canibe, N., et al. 2016: Impact of the amount of straw provided to pigs kept in intensive production conditions on the occurrence and severity of gastric ulceration at slaughter. Res. Vet. Sci., 104, 200–206. DOI: 10.1016/j. rvsc.2015.12.017
- 23. Almeida, L. R., Costa, P. S., Nascimento, A. M. A., Reis, M. de P., Barros, K. O., Alvim, L. B., et al., 2018: Porcine stomachs with and without gastric ulcer differ in Lactobacillus load and strain characteristics. Can. J. Microbiol., 64, 7, 493–499. DOI: 10.1139/cjm-2017-0758
- 24. Hellemans, A., Chiers, K., Maes, D., De Bock, M., Decostere, A., Haesebrouck, F., et al., 2011: Prevalence of “Candidatus Helicobacter suis” in pigs of different ages. Vet. Rec., 161, 6, 189–192. DOI: 10.1136/vr.161.6.189
- 25. Bosschem, I., Flahou, B., Van Deun, K., De Koker, S., Volf, J., Smet, A., et al., 2017: Species-specific immunity to Helicobacter suis. Helicobacter, 22, 3, e12375. DOI: 10.1111/hel.12375
- 26. De Witte, C., Devriendt, B., Flahou, B., Bosschem, I., Ducatelle, R., Smet, A., Haesebrouck, F. 2017: Helicobacter suis induces changes in gastric inflammation and acid secretion markers in pigs of different ages. Vet. Res., 48, 34. DOI: 10.1186/s13567-017-0441-6
- 27. De Bruyne, E., Flahou, B., Chiers, K., Meyns, T., Kumar, S., Vermoote, M., et al. 2012: An experimental Helicobacter suis infection causes gastritis and reduced daily weight gain in pigs. Vet. Microbiol., 160, 3–4, 449–454. DOI: 10.1016/j. vetmic.2012.06.031
- 28. Fabisiak, M.; Sapierzy ń ski, R.; Salamaszy ń ska-Guz, A.; Kizerwetter-Swida, M. 2010: The first description of gastric Helicobacter in freeranging wild boar (Sus scrofa) from Poland. Pol. J. Vet. Sci., 13, 1, 171–174. PMID: 21077447
- 29. Cortez Nunes, F., Letra Mateus, T., Taillieu, E., Teixeira, S., Carolino, N., Rema, A., et al., 2022: Molecular detection of Helicobacter spp. and Fusobacterium gastrosuis in pigs and wild boars and its association with gastric histopatho-logical alterations. Vet Res., 53, 78. DOI: 10.1186/s13567-022-01101-5
- 30. Guevara, B., Cogdill, A. G., 2020: Helicobacter pylori: A Review of Current Diagnostic and Management Strategies. Dig. Dis. Sci., 65, 7, 1917–1931.
- 31. Flahou, B., Rossi, M., Bakker, J., Langermans, J. A. M., Heuvelman, E., Solnick, J. V., et al. 2017: Evidence for a primate origin of zoonotic Helicobacter suis colonizing domesticated pigs. ISME J., 12, 1, 77–86. DOI: 10.1038/ismej.2017.145
- 32. Krepelkov á, Z., Karaffov á, V., B á rdov á, K., Andra š kov á, S., Novotn ý, J., 2025: Post mortem laboratory detection of Helicobacter spp. in swine. Vet. Res. Commun., 50, 15, 1–10. DOI: 10.1007/s11259-025-10961-z
- 33. De Witte, C., Demeyere, K., De Bruyckere, S., Taminiau, B., Daube, G., Ducatelle, R., et al., 2019: Characterization of the nonglandular gastric region microbiota in Helicobacter suisinfected versus noninfected pigs identifies a potential role for Fusobacterium gastrosuis in gastric ulceration. Vet. Res., 50, 39. DOI: 10.1186/s13567-019-0656-9
- 34. Krepelkov á, Z., Novotn ý, J., B á rdov á, K., Link, R., Cs ö rg ö, A., 2024: Gastric ulcers in pigs – a review. Folia Vet., 68, 2, 33–42. DOI: 10.2478/fv-2024-0015
- 35. Padra, M., Adamczyk, B., Flahou, B., Erhardsson, M., Chahal, G., Smet, A., et al., 2019: Helicobacter suis infection alters glycosylation and decreases the pathogen growthinhibiting effect and binding avidity of gastric mucins. Mucosal Immunol., 12, 3, 784–794. PMID: 30846831
- 36. Amory, J. R., Mackenzie, A. M., Pearce, G. P., 2006: Factors in the housing environment of finisher pigs associated with the development of gastric ulcers. Vet. Rec., 158, 8, 260–264. doi:10.1136/vr.158.8.260
- 37. Taillieu, E., Taelman, S., DeBruyckere, S., Goosens, E., Chantziaras, I., Steenkiste, V., et al., 2024: The role of Helicobacter suis, Fusobacterium gastrosuis, and the pars oesophageal microbiota in gastric ulceration in slaughter pigs receiving meal or pelleted feed. Vet. Res., 55, 15. DOI: 10.1186/s13567-024-01274-1
- 38. Meier, R.K., Ryser-Degiorgis, M., 2018: Wild boar and infectious diseases: Evaluation of the current risk to human and domestic animal health in Switzerland: A review. Schweiz. Arch. Tierheilkd., 160, 7–8, 443–460. DOI: 10.17236/sat00168
- 39. Mladenova-Hristova, I., Grekova, O., Patel, A., 2017: Zoonotic potential of Helicobacter spp. J. Microbiol. Immunol. Infect., 50, 3, 265–269. DOI: 10.1016/j.jmii.2016.11.003
- 40. Ball, M. E. E., Magowan, E., 2012: The effect of level of wheat inclusion in diets for growing and finishing pigs on performance, nutrient digestibility and gastric ulceration. Asian-Aust. J. Anim. Sci., 25, 7, 988–993. DOI: 10.5713/ajas.2010.10256
- 41. Reynolds, L., Rantzer, A., Friendship, R., 2012: Risk factors and management of gastric ulcers. Anim. Health Res. Rev., 13, 17–27.
- 42. Nasiadka, P., Janiszewski, P., 2015: Food preferences of wild boars (Sus scrofa L.) in the summer and early autumn expressed by the damage caused in agricultural crops. Sylwan, 159, 4, 307–317. DOI: 10.26202/sylwan.2014082
- 43. Jo, Y. Y, Choi, M. J., Chung, W. L., Hong, J. S., Lim, J. S., Kim, Y. Y., 2021: Effects of feed form and particle size on growth performance, nutrient digestibility, carcass characteristics, and gastric health in growing-finishing pigs. Anim Biosci., 34, 6, 1061–1069. DOI: 10.5713/ab.20.0777
- 44. Canibe, N., Blaabjerg, K., Lauridsen, C., 2016: Gastric ulcers in pigs. Available online: https://pure.au.dk/ws/files/108305983/Vidensyntese_Gastric_ulcers_in_pigs_231216
- 45. Friendship, R. M., 2004: Gastric ulceration in swine. J. Swine Health Prod., 12, 1, 34–35. DOI: 10.54846/jshap/381