Table 1.
Construction rules for tree-species groups. The fertility classes correspond to Orlov’s bonity categories, where 0 indicates the greatest and 6 the lowest fertility. The bonity class 3 is excluded from soil fertility-dependent subsets. Forest inventory data do not distinguish between silver birch and downy birch. However, it is known that downy birch dominates on poor soils.
| Group | Species | Share(%) | Fertility classes |
|---|---|---|---|
| All stands | All | All | all |
| Evergreen needleleaved | Norway spruce, Scots pine | >85 | all |
| Deciduous broadleaved | Norway spruce excluded, Scots pine excluded | ≥85 | all |
| Pine | Scots pine | ≥80 | all |
| Pine (fertile soil] | Scots pine | ≥ 80 | 0,1,2 |
| Pine (poor soil) | Scots pine | ≥80 | 4,5,6 |
| Birch spp. | Silver birch, downy birch | ≥80 | all |
| Birch (fertile soil) | Silver birch | ≥80 | 0,1,2 |
| Birch (poor soil) | (Silver birch), downy birch | ≥80 | 4,5,6 |
| Spruce | Norway spruce | ≥80 | all |
| Aspen | European aspen | ≥75 | all |
| Alder | Grey alder | ≥75 | all |
Table 2.
Detailed descriptions of the tree species groups introduced in Table 1. KU signifies Norway spruce, MA Scots pine, HB European aspen, KS both silver birch and downy birch (these two species are not distinguished in the forest inventory of the Estonian Environment Agency), LV grey alder, and KX species compositions not matching any of the other codes. Stands in the evergreen needleleaf group Everg. N. consist predominantly of Norway spruce and Scots pine, while stands in the group Dec. Br. are dominated by deciduous broadleaved species. In the Bon (bonity) column, code A signifies all fertility classes, P poor soils, and F fertile soils. For each tree species group, an entry in the N column gives the number of stands in that group, S its aggregate area, A the mean value of its various stand ages, H100 the mean value of the 100-year heights predicted for its various stands (this is a measure of site productivity), T its relative density, and G its basal area.
| Group | Stands | Stand structure | Tree species composition (%) | ||||||||||
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| Species | Bon | N | S (ha) | A | H100 (m) | T (%) | G (m2ha−1) | HB | KS | KU | LV | MA | KX |
| All | A | 92,230 | 231,687 | 77 | 26.6 | 94 | 28.4 | 6 | 29 | 26 | 3 | 33 | 3 |
| Everg. N. | A | 34,073 | 83,321 | 92 | 25.4 | 92 | 30.4 | 0 | 4 | 28 | 0 | 68 | 0 |
| Dec. Br. | A | 22,659 | 60,990 | 61 | 27.1 | 95 | 25.4 | 13 | 60 | 9 | 8 | 1 | 9 |
| MA | A | 1,5898 | 39,765 | 94 | 23.4 | 90 | 29.7 | 0 | 3 | 5 | 0 | 92 | 0 |
| MA | F | 8,675 | 20,553 | 85 | 27.2 | 91 | 31.1 | 0 | 3 | 7 | 0 | 90 | 0 |
| MA | P | 3,388 | 9,575 | 107 | 15.8 | 89 | 26.6 | 0 | 3 | 2 | 0 | 95 | 0 |
| KU | A | 5,092 | 11,627 | 51 | 31.3 | 93 | 27.7 | 1 | 6 | 90 | 1 | 1 | 1 |
| KS | A | 6,471 | 17,024 | 56 | 24.8 | 93 | 21.7 | 1 | 90 | 4 | 1 | 2 | 2 |
| KS | F | 4,013 | 10,130 | 50 | 27.3 | 93 | 22.6 | 1 | 90 | 5 | 1 | 1 | 2 |
| KS | P | 6,66 | 1,978 | 69 | 17.6 | 94 | 18.5 | 0 | 91 | 2 | 0 | 4 | 3 |
| HB | A | 441 | 951 | 65 | 31.7 | 95 | 31.7 | 83 | 8 | 6 | 1 | 0 | 2 |
| LV | A | 960 | 2,222 | 45 | 27.0 | 96 | 25.8 | 2 | 5 | 3 | 88 | 0 | 2 |

Figure 1.
The average seasonal course of the red ρ0.6 and near-infrared ρ2.2 reflectance ratio, summarised as a boxplot using 5-day bins. The dashed line corresponds to the preliminary model estimated from USGS Landsat-8/9 OLI L2 surface reflectance data. The wide line corresponds to model (1); the narrower line to our spline-based refinement.
Table 3.
Estimated parameters for the model (1). An observation is one stand on an image granule. RSE is the model residual error. R2 is the determination coefficient as defined by equation (2). Parameter values in the shaded cells were not significant. The parameter c has been fixed to the constant c = 1.
| Model for group | N. obs | R2 | RSE | Parameter estimates | ||||||
|---|---|---|---|---|---|---|---|---|---|---|
| a | b | d | kpk | kk | kps | ks | ||||
| All | 5.6 × 106 | 0.15 | 0.69 | 2.22 | 2.22 | 1.24 | 140.0 | 5.84 | 262.4 | 22.0 |
| Evergreen needleleaf | 2.2 × 106 | 0.02 | 0.47 | 2.21 | 39.2 | 6.52 | 140.0 | 5.03 | 270.9 | 22.0 |
| Deciduous broadleaf | 1.4 × 106 | 0.45 | 0.68 | 2.34 | 1.22 | 0.95 | 138.0 | 4.50 | 255.0 | 25.0 |
| Scots pine (MA) | 1.0 × 106 | 0.05 | 0.43 | 2.20 | 160.0 | 5.61 | 136.9 | 4.00 | 262.3 | 21.0 |
| MA (fertile soil) | 6.3 × 105 | 0.05 | 0.43 | 2.21 | 160.0 | 5.41 | 141.0 | 4.45 | 264.7 | 23.0 |
| MA (poor soil) | 6.3 × 10s | 0.04 | 0.44 | 2.17 | 99.9 | 6.83 | 140.0 | 4.61 | 261.8 | 22.0 |
| Norway spruce (KU) | 6.3 × 105 | 0.02 | 0.56 | 2.19 | 5.55 | 6.13 | 150.0 | 8.00 | 280.0 | 22.0 |
| Birch spp. (KS) | 6.3 × 105 | 0.46 | 0.63 | 2.29 | 1.23 | 0.90 | 135.0 | 4.00 | 255.0 | 25.0 |
| KS (fertile soil) | 6.3 × 105 | 0.46 | 0.64 | 2.31 | 1.25 | 0.90 | 135.0 | 4.00 | 255.0 | 25.0 |
| KS (poor soil) | 6.3 × 104 | 0.45 | 0.66 | 2.24 | 1.19 | 0.90 | 137.4 | 4.00 | 255.0 | 25.0 |
| European aspen (HB) | 3.5 × 104 | 0.47 | 0.67 | 2.35 | 1.09 | 0.90 | 138.0 | 4.20 | 250.0 | 22.0 |
| Grey alder (LV) | 4.3 × 104 | 0.49 | 0.61 | 2.36 | 0.82 | 0.90 | 131.0 | 9.27 | 257.5 | 18.9 |

Figure 2.
Evergreen needleleaf tree species group. Figure details are as in Figure 1.

Figure 3.
Deciduous broadleaved tree species group. Figure details are as in Figure 1.

Figure A1.1.
Scots pine stands.

Figure A1.2.
Scots pine stands on fertile soils.

Figure A1.3.
Scots pine stands on poor soils.

Figure A1.4.
Norway spruce stands.

Figure A1.5.
Birch stands.

Figure A1.6.
Birch stands on fertile soils.

Figure A1.7.
Birch stands on poor soils.

Figure A1.8.
European aspen stands.

Figure A1.9.
Grey alder stands.
Table A2.1.
Smoothed course of the near-infrared/red reflectance ratio ρ2.2/ρ0.6. Values are scaled by 100. Group descriptions are given in Tables 1 and 2.
| Day | Tree species groups | |||||||||||
|---|---|---|---|---|---|---|---|---|---|---|---|---|
| All | Everg. N. | Dec. Br. | MA-A | MA-F | MA-P | KU-A | KS-A | KS-F | KS-P | HB-A | LV-A | |
| 70 | 226 | 223 | 226 | 224 | 228 | 220 | 224 | 225 | 222 | 229 | 233 | 246 |
| 73 | 226 | 223 | 228 | 223 | 227 | 220 | 225 | 229 | 227 | 229 | 233 | 246 |
| 76 | 226 | 222 | 230 | 222 | 226 | 219 | 225 | 231 | 231 | 228 | 233 | 245 |
| 79 | 225 | 222 | 231 | 222 | 225 | 218 | 225 | 232 | 232 | 226 | 232 | 245 |
| 82 | 224 | 222 | 230 | 221 | 224 | 218 | 224 | 231 | 232 | 224 | 231 | 245 |
| 85 | 223 | 222 | 229 | 221 | 223 | 217 | 222 | 229 | 229 | 223 | 231 | 244 |
| 88 | 222 | 222 | 227 | 220 | 222 | 216 | 220 | 226 | 226 | 221 | 230 | 244 |
| 91 | 222 | 222 | 226 | 220 | 222 | 216 | 219 | 224 | 224 | 221 | 230 | 243 |
| 94 | 221 | 222 | 227 | 220 | 221 | 216 | 218 | 224 | 224 | 223 | 230 | 243 |
| 97 | 221 | 222 | 228 | 220 | 221 | 216 | 219 | 226 | 226 | 224 | 231 | 243 |
| 100 | 221 | 222 | 230 | 220 | 221 | 217 | 220 | 229 | 229 | 226 | 232 | 243 |
| 103 | 222 | 222 | 232 | 220 | 221 | 217 | 220 | 232 | 232 | 227 | 233 | 244 |
| 106 | 222 | 222 | 232 | 220 | 221 | 217 | 220 | 233 | 233 | 227 | 233 | 245 |
| 109 | 222 | 222 | 231 | 220 | 221 | 217 | 219 | 231 | 231 | 226 | 232 | 246 |
| 112 | 223 | 222 | 230 | 220 | 222 | 218 | 219 | 230 | 230 | 224 | 231 | 248 |
| 115 | 223 | 222 | 229 | 221 | 222 | 218 | 219 | 229 | 229 | 222 | 229 | 251 |
| 118 | 225 | 222 | 229 | 221 | 222 | 219 | 220 | 228 | 229 | 222 | 228 | 254 |
| 121 | 227 | 222 | 231 | 221 | 223 | 219 | 222 | 230 | 231 | 222 | 229 | 259 |
| 124 | 229 | 222 | 234 | 221 | 223 | 219 | 224 | 234 | 236 | 224 | 233 | 265 |
| 127 | 232 | 222 | 240 | 221 | 223 | 220 | 225 | 239 | 242 | 228 | 239 | 272 |
| 130 | 236 | 222 | 250 | 221 | 223 | 219 | 226 | 248 | 252 | 235 | 250 | 280 |
| 133 | 241 | 222 | 265 | 221 | 222 | 219 | 227 | 262 | 267 | 246 | 266 | 289 |
| 136 | 245 | 222 | 281 | 221 | 222 | 219 | 229 | 278 | 283 | 261 | 285 | 298 |
| 139 | 250 | 223 | 296 | 220 | 221 | 218 | 231 | 293 | 297 | 275 | 302 | 308 |
| 142 | 253 | 223 | 307 | 220 | 220 | 217 | 232 | 304 | 307 | 287 | 316 | 316 |
| 145 | 256 | 223 | 314 | 219 | 220 | 217 | 233 | 311 | 313 | 293 | 326 | 324 |
| 148 | 258 | 223 | 316 | 219 | 220 | 217 | 234 | 312 | 314 | 296 | 330 | 330 |
| 151 | 260 | 223 | 314 | 219 | 220 | 217 | 234 | 308 | 310 | 296 | 330 | 336 |
| 154 | 260 | 223 | 312 | 219 | 220 | 217 | 233 | 304 | 307 | 295 | 329 | 340 |
| 157 | 261 | 223 | 312 | 219 | 220 | 217 | 232 | 304 | 306 | 297 | 330 | 343 |
| 160 | 261 | 223 | 314 | 219 | 220 | 217 | 232 | 309 | 310 | 302 | 331 | 345 |
| 163 | 262 | 223 | 318 | 219 | 220 | 217 | 233 | 315 | 316 | 308 | 332 | 347 |
| 166 | 262 | 223 | 321 | 219 | 220 | 217 | 234 | 319 | 320 | 312 | 333 | 349 |
| 169 | 262 | 223 | 320 | 219 | 220 | 217 | 234 | 317 | 318 | 311 | 332 | 350 |
| 172 | 262 | 223 | 318 | 218 | 219 | 217 | 234 | 312 | 312 | 308 | 330 | 351 |
| 175 | 262 | 223 | 315 | 218 | 219 | 216 | 234 | 307 | 308 | 304 | 328 | 352 |
| 178 | 262 | 223 | 313 | 218 | 219 | 216 | 233 | 304 | 305 | 303 | 327 | 353 |
| 181 | 262 | 223 | 313 | 218 | 218 | 215 | 233 | 306 | 307 | 304 | 326 | 354 |
| 184 | 263 | 223 | 315 | 218 | 218 | 215 | 232 | 312 | 313 | 307 | 326 | 356 |
| 187 | 264 | 223 | 317 | 218 | 218 | 215 | 232 | 317 | 318 | 310 | 326 | 357 |
| 190 | 264 | 223 | 318 | 219 | 219 | 216 | 232 | 318 | 320 | 310 | 325 | 359 |
| 193 | 266 | 222 | 318 | 219 | 219 | 216 | 234 | 315 | 318 | 307 | 324 | 361 |
| 196 | 267 | 222 | 318 | 219 | 220 | 217 | 236 | 311 | 314 | 304 | 323 | 363 |
| 199 | 268 | 222 | 319 | 220 | 220 | 218 | 238 | 310 | 312 | 302 | 323 | 364 |
| 202 | 269 | 222 | 320 | 220 | 221 | 219 | 241 | 311 | 313 | 302 | 324 | 365 |
| 205 | 269 | 222 | 322 | 220 | 222 | 219 | 243 | 312 | 314 | 304 | 326 | 364 |
| 208 | 269 | 221 | 323 | 220 | 222 | 220 | 245 | 314 | 315 | 307 | 328 | 363 |
| 211 | 269 | 221 | 324 | 220 | 222 | 220 | 247 | 317 | 317 | 311 | 328 | 362 |
| 214 | 268 | 220 | 325 | 220 | 222 | 221 | 247 | 320 | 319 | 315 | 328 | 360 |
| 217 | 267 | 220 | 323 | 219 | 222 | 221 | 246 | 320 | 319 | 316 | 325 | 357 |
| 220 | 265 | 219 | 320 | 219 | 221 | 220 | 244 | 316 | 315 | 313 | 321 | 353 |
| 223 | 262 | 218 | 315 | 218 | 220 | 219 | 242 | 311 | 311 | 309 | 317 | 349 |
| 226 | 259 | 218 | 311 | 216 | 219 | 218 | 240 | 305 | 306 | 302 | 312 | 345 |
| 229 | 255 | 217 | 306 | 215 | 217 | 216 | 237 | 300 | 302 | 294 | 307 | 341 |
| 232 | 251 | 216 | 299 | 213 | 215 | 214 | 234 | 293 | 296 | 285 | 301 | 336 |
| 235 | 247 | 215 | 291 | 211 | 213 | 211 | 231 | 283 | 286 | 276 | 294 | 331 |
| 238 | 242 | 214 | 282 | 209 | 211 | 208 | 228 | 274 | 276 | 269 | 287 | 326 |
| 241 | 239 | 214 | 275 | 208 | 209 | 206 | 226 | 266 | 267 | 263 | 282 | 321 |
| 244 | 235 | 213 | 269 | 206 | 207 | 203 | 224 | 261 | 261 | 258 | 278 | 316 |
| 247 | 232 | 212 | 264 | 204 | 206 | 201 | 224 | 257 | 256 | 255 | 275 | 312 |
| 250 | 230 | 212 | 259 | 203 | 205 | 199 | 224 | 252 | 251 | 252 | 273 | 308 |
| 253 | 228 | 211 | 256 | 202 | 204 | 198 | 225 | 249 | 249 | 249 | 271 | 305 |
| 256 | 226 | 211 | 254 | 201 | 203 | 197 | 227 | 247 | 247 | 246 | 270 | 303 |
| 259 | 224 | 211 | 253 | 200 | 202 | 197 | 229 | 244 | 245 | 243 | 268 | 301 |
| 262 | 223 | 211 | 252 | 200 | 201 | 197 | 230 | 242 | 242 | 239 | 265 | 300 |
| 265 | 221 | 211 | 250 | 200 | 200 | 197 | 231 | 239 | 239 | 236 | 258 | 298 |
| 268 | 219 | 211 | 246 | 200 | 200 | 199 | 232 | 236 | 237 | 233 | 248 | 296 |
| 271 | 216 | 211 | 240 | 201 | 201 | 201 | 233 | 234 | 233 | 231 | 235 | 292 |
| 274 | 213 | 211 | 230 | 203 | 202 | 203 | 232 | 228 | 227 | 227 | 220 | 288 |
| 277 | 209 | 212 | 217 | 204 | 203 | 206 | 228 | 216 | 213 | 221 | 204 | 282 |
| 280 | 205 | 212 | 201 | 206 | 205 | 209 | 219 | 199 | 193 | 212 | 189 | 276 |
| 283 | 202 | 213 | 186 | 208 | 206 | 212 | 210 | 182 | 174 | 204 | 177 | 268 |
| 286 | 199 | 213 | 176 | 211 | 209 | 215 | 204 | 171 | 161 | 199 | 171 | 260 |
| 289 | 198 | 214 | 173 | 214 | 212 | 219 | 202 | 170 | 159 | 200 | 170 | 253 |
| 292 | 200 | 215 | 177 | 217 | 216 | 224 | 206 | 179 | 166 | 206 | 175 | 245 |
| 295 | 202 | 216 | 184 | 221 | 221 | 230 | 213 | 192 | 178 | 215 | 183 | 237 |
| 298 | 207 | 217 | 194 | 225 | 227 | 236 | 225 | 204 | 190 | 224 | 193 | 230 |
| 301 | 212 | 218 | 205 | 229 | 233 | 243 | 239 | 215 | 202 | 233 | 204 | 222 |
| 304 | 217 | 218 | 216 | 233 | 239 | 250 | 254 | 225 | 212 | 241 | 215 | 215 |
| 307 | 223 | 219 | 227 | 238 | 246 | 257 | 269 | 235 | 223 | 249 | 226 | 208 |
| 310 | 228 | 220 | 238 | 242 | 252 | 263 | 284 | 245 | 234 | 257 | 237 | 200 |
Table A3.1.
The normalised difference vegetation index INDVI = (ρ0.85 – ρ0.6) / (ρ0.85 + ρ0.6) mean values and standard deviation for our species groups. Spectral reflectances are from the bands at 0.85 μm (ρ0.85) and 0.6 μm (ρ0.6). Group descriptions are given in Tables 1 and 2. The estimate is made for the period starting from day 141 and ending with day 259.
| Tree species group | NDVI | ||
|---|---|---|---|
| Species | Bonity | Mean | Standard deviation |
| All | A | 0.79 | 0.09 |
| Everg. N. | A | 0.78 | 0.06 |
| Dec. Br. | A | 0.78 | 0.13 |
| MA | A | 0.77 | 0.06 |
| MA | F | 0.78 | 0.06 |
| MA | P | 0.76 | 0.06 |
| KU | A | 0.82 | 0.07 |
| KS | A | 0.77 | 0.13 |
| KS | F | 0.78 | 0.13 |
| KS | P | 0.75 | 0.15 |
| HB | A | 0.78 | 0.15 |
| LV | A | 0.79 | 0.14 |