
Figure 1.
Locations of ENFRP permanent sample plots.
Table 1.
Distribution of cohorts by canopy layer and tree species (the number of cohorts that include at least one dominant tree height-diameter measurement pair is presented in brackets).
| Upper canopy layer | Secondary canopy layer | Undergrowth | Advanced regeneration | |
|---|---|---|---|---|
| Pinus sylvestris L. | 2077 (1968) | 10 | 0 | 0 |
| Picea abies (L.) H. Karst. | 1039 (940) | 697 | 0 | 47 |
| Betula spp. | 993 (706) | 58 | 0 | 1 |
| Populus tremula L. | 157(144) | 2 | 0 | 0 |
| Alnus glutinosa (L.) Gaertn. | 81 (49) | 0 | 0 | 0 |
| Alnus incana (L.) Moench | 60(15) | 1 | 0 | 0 |
| Salix spp. | 22 (8) | 0 | 0 | 0 |
| Fraxinus excelsior L. | 5(5) | 5 | 0 | 0 |
| Tilia cordata Mill. | 2 | 1 | 0 | 0 |
| Acer platanoides L. | 2 | 8 | 0 | 1 |
| Quercus robur L. | 1(1) | 0 | 0 | 0 |
| Sorbus aucuparia L. | 0 | 0 | 1 | 0 |
| Corylus avellana L. | 0 | 0 | 7 | 0 |
Table 2.
The standardized height-diameter equations evaluated in the study (h – tree height (m); d – tree diameter at breast height (cm); a, b – estimated parameters; c – parameter fixed as constant).
| No | Equation | Linear form | References | Decreasing region | Height at zero diameter | Height asymptote | Diameter at the inflection point |
|---|---|---|---|---|---|---|---|
| Fl | (Hoßfeld, 1823; Kiviste et al., 2002) | a<=0 | b<=0 | 1.3 | ||||
| Fla | (Hoßfeld, 1823; Kiviste et al., 2002) | a<=0 | b<=0 | 1.3 | ||||
| F2 | (Näslund, 1936) | a<=0 | b<=0 | 1.3 | ||||
| F3 | h =1.3 + adb | ln(h – 1.3) = ln a + b ln d | (Curtis, 1967) | a<=0 | b<=0 |b>l | 1.3 | – | – |
| F4 | h = 1.3 + aeb/d | (Curtis, 1967) | a<=0 | b>=0 | 1.3 | 1.3 + a | ||
| F5 | (Curtis, 1967) | a<=0 | b<=0 | 1.3 | 1.3 + a | |||
| F5a | (Mehtätalo et al., 2015) | a<=0 | b<=0 | 1.3 | – | – | ||
| F6 | (Wykoff et al., 1982) | b>=0 | 1.3 + e a+b | 1.3+ ea | |||
| F7 | h= 1.3 + ade−bd | (Lebedev, 2020) | a<=0 | b<=0 | d>l/b | 1.3 | 1.3 | ||
| F8 | h= 1.3 + a(ln(l + d))d | ln(h – 1.3) = ln a + b ln(ln(l + d)) | (Lebedev, 2020) | a<=0 | b<=0 | 1.3 | – | – |
| F9 | h =1.3 + a(l – e−bd)c | – | (Chapman, 1961; Meyer, 1940; Nigul et al., 2021; Richards, 1959) | a<=0 | b<=0 | 1.3 | 1.3 + a | |
| F10 | h = a + b log d | - | (Curtis, 1967) | b<=0 | – ∞ | – | – |
| F11 | – | (Curtis, 1967) | a<=0 | b>=0 | –∞ | a | – | |
| F12 | h = a + bd | – | Line | b<=0 | α | – | – |
| F13 | – | Hyperbole | a<=0 | b>=0 | –∞ | a | – | |
| F14 | – | Square root | a<=0|b<=0 | 0 | – | – |
Table 3.
Goodness of fit statistics of the two-parameter height-diameter curves used in the study. The meanings of the column headings are presented in Table 4.
| No | Noc | c | Nfail | Ndecr | Nlow | Nhd | RMSE | MAD | ME | ME5% | MEdom | MEDdom | a | b |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| Fl | 1 | 1.3 | 0 | 68 | 0 | 127034 | 1.406 | 1.202 | -0.0043 | 0.0034 | -0.0805 | -0.0796 | 0.7829 | 0.0365 |
| Fl | 2 | 1.5 | 1 | 33 | 0 | 127034 | 1.401 | 1.198 | -0.0013 | 0.0070 | -0.0312 | -0.0261 | 1.1954 | 0.0393 |
| Fl | 3 | 1.5117 | 1 | 33 | 0 | 127023 | 1.401 | 1.198 | -0.0011 | 0.0072 | -0.0282 | -0.0231 | 1.2262 | 0.0395 |
| Fl | 4 | 1.58 | 1 | 30 | 0 | 127034 | 1.401 | 1.198 | -0.0001 | 0.0085 | -0.0108 | -0.0033 | 1.4226 | 0.0404 |
| Fl | 5 | 1.5896 | 0 | 29 | 0 | 127034 | 1.401 | 1.197 | 0.0000 | 0.0086 | -0.0084 | -0.0013 | 1.4519 | 0.0405 |
| Fl | 6 | 1.59 | 0 | 29 | 0 | 127034 | 1.400 | 1.197 | 0.0000 | 0.0086 | -0.0083 | -0.0013 | 1.4532 | 0.0405 |
| Fl | 7 | 1.6 | 0 | 29 | 0 | 127034 | 1.400 | 1.197 | 0.0002 | 0.0088 | -0.0057 | 0.0008 | 1.4855 | 0.0407 |
| Fl | 8 | 1.6118 | 0 | 29 | 0 | 127034 | 1.400 | 1.197 | 0.0004 | 0.0090 | -0.0027 | 0.0035 | 1.5250 | 0.0408 |
| Fl | 9 | 1.6655 | 0 | 28 | 0 | 127034 | 1.400 | 1.197 | 0.0012 | 0.0100 | 0.0112 | 0.0187 | 1.7148 | 0.0414 |
| Fl | 10 | 1.7 | 0 | 28 | 0 | 127034 | 1.400 | 1.197 | 0.0017 | 0.0107 | 0.0202 | 0.0277 | 1.8505 | 0.0417 |
| Fl | 11 | 1.7128 | 0 | 28 | 0 | 127034 | 1.400 | 1.197 | 0.0019 | 0.0109 | 0.0235 | 0.0318 | 1.9038 | 0.0419 |
| Fl | 12 | 2 | 0 | 22 | 0 | 127034 | 1.405 | 1.200 | 0.0064 | 0.0161 | 0.0998 | 0.1067 | 3.5648 | 0.0444 |
| F1a | 13 | 2 | 0 | 22 | 0 | 127034 | 1.405 | 1.200 | 0.0064 | 0.0161 | 0.0998 | 0.1067 | 3.5648 | 0.0444 |
| Fl | 14 | 3 | 1 | 27 | 6 | 127024 | 1.460 | 1.260 | 0.0220 | 0.0313 | 0.3722 | 0.3642 | 35.725 | 0.0496 |
| F2 | 15 | 1 | 1 | 304 | 0 | 127025 | 1.419 | 1.221 | -0.0084 | -0.0013 | -0.1499 | -0.1605 | 0.4339 | 0.0308 |
| F2 | 16 | 2 | 0 | 22 | 0 | 127034 | 1.405 | 1.202 | -0.0019 | 0.0062 | -0.1022 | -0.1036 | 0.8874 | 0.1844 |
| F2 | 17 | 3 | 0 | 21 | 0 | 127034 | 1.404 | 1.201 | 0.0000 | 0.0086 | -0.0838 | -0.0808 | 0.9421 | 0.3274 |
| F2 | 18 | 4 | 0 | 21 | 0 | 127034 | 1.403 | 1.199 | 0.0009 | 0.0097 | -0.0742 | -0.0685 | 0.8905 | 0.4343 |
| F2 | 19 | 5 | 1 | 21 | 0 | 127025 | 1.403 | 1.200 | 0.0015 | 0.0104 | -0.0682 | -0.0610 | 0.8180 | 0.5140 |
| F2 | 20 | 10 | 4 | 21 | 0 | 126925 | 1.404 | 1.203 | 0.0025 | 0.0115 | -0.0560 | -0.0458 | 0.5417 | 0.7183 |
| F2 | 21 | 20 | 7 | 19 | 0 | 126909 | 1.405 | 1.205 | 0.0029 | 0.0121 | -0.0498 | -0.0386 | 0.3109 | 0.8479 |
| F2 | 22 | 200 | 91 | 14 | 0 | 125759 | 1.404 | 1.204 | 0.0034 | 0.0124 | -0.0450 | -0.0332 | 0.0354 | 0.9837 |
| F2 | 23 | 1000 | 1030 | 2 | 0 | 108218 | 1.389 | 1.192 | 0.0040 | 0.0132 | -0.0458 | -0.0333 | 0.0073 | 0.9967 |
| F2 | 24 | 2000 | 2055 | 0 | 0 | 85463 | 1.372 | 1.178 | 0.0049 | 0.0140 | -0.0517 | -0.0208 | 0.0036 | 0.9984 |
| F3 | 25 | – | 4 | 310 | 0 | 126977 | 1.448 | 1.251 | -0.0062 | 0.0022 | -0.2843 | -0.2888 | 4.8087 | 0.4344 |
| F4 | 26 | – | 8 | 21 | 0 | 126839 | 1.406 | 1.206 | 0.0034 | 0.0126 | -0.0434 | -0.0319 | 26.941 | -7.1578 |
| F5 | 27 | 1 | 3 | 21 | 0 | 126849 | 1.405 | 1.204 | 0.0022 | 0.0113 | -0.0595 | -0.0519 | 27.433 | 7.6564 |
| F5 | 28 | 1 | 1 | 21 | 57 | 127025 | 1.407 | 1.206 | 0.0029 | 0.0119 | -0.0477 | -0.0362 | 28.308 | 7.0243 |
| F3 | 29 | – | 2 | 125 | 8 | 127007 | 1.443 | 1.243 | -0.0044 | 0.0040 | -0.2785 | -0.2835 | 5.6670 | 0.4007 |
| F4 | 30 | – | 6 | 21 | 100 | 126864 | 1.409 | 1.212 | 0.0038 | 0.0130 | -0.0311 | -0.0194 | 27.867 | -6.5447 |
| F6 | 31 | 1 | 0 | 21 | 0 | 127034 | 1.404 | 1.200 | 0.0012 | 0.0100 | -0.0752 | -0.0727 | 3.3291 | -8.1500 |
| F5 | 32 | 1.1 | 3 | 21 | 0 | 126849 | 1.405 | 1.204 | 0.0021 | 0.0112 | -0.0609 | -0.0540 | 27.468 | 7.0015 |
| F5 | 33 | 1.3 | 2 | 21 | 0 | 126986 | 1.404 | 1.201 | 0.0020 | 0.0110 | -0.0638 | -0.0574 | 27.536 | 5.9887 |
| F5 | 34 | 8 | 0 | 20 | 0 | 127034 | 1.407 | 1.204 | -0.0012 | 0.0069 | -0.1285 | -0.1388 | 30.173 | 1.3628 |
| F5 | 35 | 0.5 | 6 | 21 | 0 | 126758 | 1.406 | 1.206 | 0.0027 | 0.0119 | -0.0518 | -0.0411 | 27.192 | 14.808 |
| Fl | 36 | 1.59 | 1 | 21 | 0 | 127034 | 1.400 | 1.197 | 0.0000 | 0.0086 | -0.0083 | -0.0013 | 17.143 | 0.2572 |
| Fl | 37 | 1.6 | 0 | 22 | 0 | 127034 | 1.400 | 1.197 | 0.0002 | 0.0088 | -0.0057 | 0.0008 | 17.145 | 0.2553 |
| F5a | 38 | 1 | 12 | 285 | 0 | 126822 | 1.444 | 1.247 | -0.0071 | 0.0011 | -0.2627 | -0.2675 | 5.5728 | 0.6048 |
| F7 | 39 | – | 2 | 282 | 0 | 127013 | 1.414 | 1.213 | -0.0045 | 0.0024 | 0.0617 | 0.0654 | 1.7051 | 0.0293 |
| F8 | 40 | – | 3 | 20 | 0 | 126997 | 1.426 | 1.230 | -0.0040 | 0.0040 | -0.2284 | -0.2337 | 4.0636 | 1.3151 |
| F9 | 41 | 1 | 327 | 187 | 0 | 121663 | 1.417 | 1.220 | -0.0075 | 0.0004 | -0.0475 | -0.0516 | 23.967 | 0.0839 |
| F9 | 42 | 0.80312 | 856 | 23 | 0 | 109582 | 1.410 | 1.216 | -0.0062 | 0.0033 | -0.0946 | -0.1033 | 24.840 | 0.0669 |
| F9 | 42 | 1.18444 | 242 | 66 | 0 | 123709 | 1.410 | 1.210 | -0.0049 | 0.0028 | -0.0023 | -0.0064 | 23.109 | 0.0988 |
| F9 | 44 | 1.19374 | 230 | 65 | 0 | 123885 | 1.409 | 1.210 | -0.0048 | 0.0029 | -0.0001 | -0.0039 | 23.090 | 0.0993 |
| F9 | 45 | 1.19 | 233 | 67 | 0 | 123846 | 1.409 | 1.210 | -0.0048 | 0.0029 | -0.0010 | -0.0050 | 23.102 | 0.0991 |
| F10 | 46 | – | 0 | 3262 | 55 | 127034 | 1.420 | 1.221 | 0.0000 | 0.0080 | -0.1941 | -0.1922 | -1.5402 | 7.1452 |
| F1l | 47 | – | 0 | 21 | 226 | 127034 | 1.590 | 1.340 | 0.0000 | -0.0015 | 0.4462 | 0.4068 | 21.162 | -752.43 |
| F12 | 48 | – | 0 | 136 | ü | 127034 | 1.493 | 1.289 | 0.0000 | 0.0118 | -0.4066 | -0.4012 | 10.427 | 0.4127 |
| F13 | 49 | – | 0 | 21 | 172 | 127034 | 1.466 | 1.249 | 0.0000 | 0.0046 | 0.1151 | 0.1165 | 25.051 | -108.27 |
| F14 | 50 | – | 0 | 1585 | 11 | 127034 | 1.443 | 1.245 | 0.0000 | 0.0090 | -0.3161 | -0.3112 | 3.2249 | 3.4302 |
Table 4.
The meanings of the column headings of Table 3.
| Meaning | |
|---|---|
| No | Height-diameter equation number (as presented in Table 2) |
| Noc | Height-diameter curve number |
| Nfail | Number of cohorts for which the non-linear regression did not converge. |
| Ndecr | Number of cohorts for which the estimates for parameters a and b resulted in the height-diameter curve decreasing. |
| Nlow | Number of height-diameter measurement pairs for which the model predicted a height lower than 1.3 m. |
| The following characteristics were calculated from the trimmed dataset, from which cohorts with extreme distributions were excluded: | |
|---|---|
| Nhd | The number of height-diameter measurement pairs at the individual tree level. |
| RMSE | The root mean square error of height prediction. |
| MAD | The median absolute error of height prediction. |
| ME | The mean error based of height prediction. |
| ME5% | The trimmed mean error of height prediction error (excluding the largest and smallest 5% of the data). |
| MEdom | The mean height prediction error based on dominant tree measurement data. |
| MEDdom | The median value of height prediction error based on dominant tree measurement data. |
| a | The median values of parameter a, estimated for cohorts with non-linear regression |
| b | The median values of parameter b, estimated for cohorts with non-linear regression |

Figure 2.
Median errors of dominant tree height measurement predictions by cohort mean diameter classes for each out of 15 selected height-diameter curves. The numbers of dominant tree height measurements by mean diameter classes are presented at the bottom of the plots. The height-diameter curve numbers (Noc in Table 3) in the legend are in ascending order of the absolute value of the median prediction error of dominant height.