
Evaluating Livestock Culling Practices Using Population Projection Models in Neolithic Dalmatia
By: Nicholas Triozzi and Sarah B. McClure
References
- Awemu, EM, Nwakalor, LN and Abubakar, BY. 1999. ‘Environmental influences on preweaning mortality and reproductive performance of Red Sokoto does’. Small Rumin. Res., 34: 161–165. DOI: 10.1016/S0921-4488(99)00058-9
- Balasse, M, Chemineau, P, Parisot, S, Fiorillo, D and Keller, M. 2023. ‘Experimental Data from Lacaune and Merino Sheep Provide New Methodological and Theoretical Grounds to Investigate Autumn Lambing in Past Husbandries’. J. Archaeol. Method Theory. DOI: 10.1007/s10816-022-09600-7
- Bogaard, A, Arbogast, R-M, Ebersbach, R, Fraser, RA, Knipper, C, Krahn, C, Schäfer, M, Styring, AK and Krause, R. 2016. The Bandkeramik settlement of Vaihingen an der Enz, Kreis Ludwigsburg (Baden-Württemberg): an integrated perspective on land use, economy and diet.
- Bradburd, D. 1982. ‘Volatility of Animal Wealth among Southwest Asian Pastoralists’. Hum. Ecol., 10: 85–106. DOI: 10.1007/BF01531106
- Bradburd, DA. 1980. ‘Never Give a Shepherd an Even Break: Class and Labor among the Komachi’. Am. Ethnol., 7: 603–620. DOI: 10.1525/ae.1980.7.4.02a00010
- Brochier, JÉ. 2013. ‘The use and abuse of culling profiles in recent zooarchaeological studies: Some methodological comments on “frequency correction” and its consequences’. J. Archaeol. Sci., 40: 1416–1420. DOI: 10.1016/j.jas.2012.09.028
- Browman, DL. 1987. ‘Agro-pastoral risk management in the Central Andes’. Res. Econ. Anthropol., 8: 171–200.
- Browman, DL. 1997. ‘Pastoral Risk Perception and Risk Definition for Altiplano Herders’. Nomad. People., 1: 22–36. DOI: 10.3167/082279497782384695
- Buckley, M, Whitcher Kansa, S, Howard, S, Campbell, S, Thomas-Oates, J and Collins, M. 2010. ‘Distinguishing between archaeological sheep and goat bones using a single collagen peptide’. J. Archaeol. Sci., 37: 13–20. DOI: 10.1016/j.jas.2009.08.020
- Crouse, DT, Crowder, LB and Caswell, H. 1987. ‘A Stage-Based Population Model for Loggerhead Sea Turtles and Implications for Conservation’. Ecology, 68: 1412–1423. DOI: 10.2307/1939225
- Dahl, G and Hjort, A. 1976. Having herds: pastoral herd growth and household economy, Dept. of Social Anthropology. University of Stockholm, Stockholm.
- Debele, G and Duguma, M. 2013. ‘Study on major causes of kid mortality in Adami Tulu Jido Kombolcha District of Oromia, Ethiopia’. Agric. Biol. J. North Am., 4: 110–115. DOI: 10.5251/abjna.2013.4.2.110.115
- Debele, G, Duguma, M and Hundessa, F. 2011. ‘Effect of different factors on mortality rate of Arsi- Bale kids in mid rift valley of Ethiopia’. Glob. Vet., 6: 56–60.
- Diekmann, Y, Gillis, RE, Lu, Z, Rudzinski, A, De Iorio, M and Thomas, MG. 2026. ‘Bayesian Inference of Sex-Specific Mortality Profiles and Product Yields from Unsexed Cattle Zooarchaeological Remains’. J. Archaeol. Method Theory, 33:
13 . DOI: 10.1007/s10816-025-09749-x - Dyson-Hudson, R and Dyson-Hudson, N. 1969. ‘Subsistence Herding in Uganda’. Sci. Am., 220: 76–89. DOI: 10.1038/scientificamerican0269-76
- Erasmus, JA, Fourie, AJ and Venter, JJ. 1985. ‘Influence of age on reproductive performance of the Improved Boer goat doe’. S. Afr. J. Anim. Sci., 15: 5–7.
- Fabre, M, Forest, V, Ranché, C, Fiorillo, D, Casabianca, F, Vigne, J-D and Balasse, M. 2023. ‘Milk and meat exploitation, autumn lambing and use of forest resources in Neolithic Corsican sheep farming systems (fifth to third millennia cal BC)’. J. Archaeol. Sci. Reports, 50:
104037 . DOI: 10.1016/j.jasrep.2023.104037 - Galina, MA, Silva, E, Morales, R and Lopez, B. 1995. ‘Reproductive performance of Mexican dairy goats under various management systems’. Small Rumin. Res., 18: 249–253. DOI: 10.1016/0921-4488(95)00693-6
- Galvin, K. 1985. Food procurement, diet and nutrition of Turkana pastoralists in an ecological and social context, State University of New York at Binghamton PP – United States — New York, United States — New York.
- Hadjikoumis, A. 2017.
‘Age-at-death in traditional Cypriot sheep and goat husbandry’ . In: Rowley-Conwy, P, Serjeanston, D and Halstead, P (eds.) Economic Zooarchaeology. Oxbow Books. DOI: 10.2307/j.ctt1s4751q.20 - Halstead, P. 1996. ‘Pastoralism or Household Herding? Problems of Scale and Specialization in Early Greek Animal Husbandry’. World Archaeol., 28: 20–42. DOI: 10.1080/00438243.1996.9980329
- Halstead, P. 1998a. ‘Ask the Fellows who Lop the Hay Leaf-Fodder in the Mountains of Northwest Greece’. Rural Hist., 9: 211–234. DOI: 10.1017/S0956793300001588
- Halstead, P. 1998b. ‘Mortality models and milking: problems of uniformitarianism, optimality and equifinality reconsidered’. Anthropozoologica, 27: 3–20.
- Halstead, P. 2024. ‘Zooarchaeological evidence for livestock management in (earlier) Neolithic Europe: Outstanding questions and some limitations of current approaches’. Quat. Int., 683–684: 42–50. DOI: 10.1016/j.quaint.2023.09.013
- Hatcher, S, Eppleston, J, Graham, RP, McDonald, J, Schlunke, S, Watt, B and Thornberry, KJ. 2008. ‘Higher weaning weight improves postweaning growth and survival in young Merino sheep’. Aust. J. Exp. Agric., 48:
966 . DOI: 10.1071/EA07407 - Helmer, D, Gourichon, L and Vila, E. 2007. ‘The development of the exploitation of products from Capra and Ovis (meat, milk and fleece) from the PPNB to the Early Bronze in the northern Near East (8700 to 2000 BC cal.)’. Anthropozoologica, 2: 41–69.
- Henrich, J and McElreath, R. 2002. ‘Are Peasants Risk-Averse Decision Makers?’ Curr. Anthropol., 43: 172–181. DOI: 10.1086/338291
- Kuznar, LA. 1991. ‘Transhumant Goat Pastoralism in the High Sierra of the South Central Andes: Human Responses to Environmental and Social Uncertainty’. Nomad. People, 93–104.
- Kuznar, LA. 2001. ‘Risk Sensitivity and Value among Andean Pastoralists: Measures, Models, and Empirical Tests’. Curr. Anthropol., 42: 432–440. DOI: 10.1086/320483
- Lefkovitch, LP. 1965. ‘The study of population growth in organisms grouped by stages’. Biometrics, 21: 1–18. DOI: 10.2307/2528348
- Lesnoff, M. 2024. mmage: A R package for sex-and-age population matrix models.
- Lesnoff, M, Lancelot, R, Tillard, E and Dohoo, IR. 2000. ‘A steady-state approach of benefit–cost analysis with a periodic Leslie-matrix model’. Prev. Vet. Med., 46: 113–128. DOI: 10.1016/S0167-5877(00)00139-2
- Mace, R. 1993. ‘Nomadic Pastoralists Adopt Subsistence Strategies That Maximise Long-Term Household Survival’. Behav. Ecol. Sociobiol., 33: 329–334. DOI: 10.1007/BF00172931
- Malher, X, Seegers, H and Beaudeau, F. 2001. ‘Culling and mortality in large dairy goat herds managed under intensive conditions in western France’. Livest. Prod. Sci., 71: 75–86. DOI: 10.1016/S0301-6226(01)00242-1
- Marković, M. 1987. ‘Stočarstvo na Dinari’. Stočarstvo, 41: 257–273.
- Marom, N and Bar-Oz, G. 2009. ‘Culling profiles: the indeterminacy of archaeozoological data to survivorship curve modelling of sheep and goat herd maintenance strategies’. J. Archaeol. Sci., 36: 1184–1187. DOI: 10.1016/j.jas.2009.01.007
- Mazzucco, N, Guilbeau, D, Kačar, S, Podrug, E, Forenbaher, S, Radić, D and Moore, AMT. 2018. ‘The time is ripe for a change. The evolution of harvesting technologies in Central Dalmatia during the Neolithic period (6th millennium cal BC)’. J. Anthropol. Archaeol., 51: 88–103. DOI: 10.1016/j.jaa.2018.06.003
- McClure, SB, Magill, C, Podrug, E, Moore, AMT, Harper, TK, Culleton, BJ, Kennett, DJ and Freeman, KH. 2018. ‘Fatty acid specific δ 13 C values reveal earliest Mediterranean cheese production 7,200 years ago’. 1–15. DOI: 10.1371/journal.pone.0202807
- McClure, SB and Podrug, E. 2016.
‘Villages, Landscapes, and Early Farming in Northern Dalmatia’ . In: Lillios, KT and Chazan, M (eds.) Fresh Fields and Pastures New’, Sidestone Press, Leiden, pp. 117–144. - McClure, SB, Podrug, E, Jović, J, Monroe, S, Radde, HD, Triozzi, N, Welker, MH and Zavodny, E. 2022. ‘The Zooarchaeology of Neolithic farmers: Herding and hunting on the Dalmatian coast of Croatia’. Quat. Int., 634: 27–37. DOI: 10.1016/j.quaint.2022.06.013
- McClure, SB, Podrug, E, Moore, AMT, Culleton, BJ and Kennett, DJ. 2014. ‘AMS 14C Chronology and Ceramic Sequences of Early Farmers in the Eastern Adriatic’. Radiocarbon, 56: 1019–1038. DOI: 10.2458/56.17918
- Mellado, M, Valdéz, R, García, JE, López, R and Rodríguez, A. 2006. ‘Factors affecting the reproductive performance of goats under intensive conditions in a hot arid environment’. Small Rumin. Res., 63: 110–118. DOI: 10.1016/j.smallrumres.2005.02.016
- Moritz, M, Buffington, A, Yoak, AJ, Hamilton, IM and Garabed, R. 2017. ‘No Magic Number: an Examination of the Herd-Size Threshold in Pastoral Systems Using Agent-Based Modeling’. Hum. Ecol., 45: 525–532. DOI: 10.1007/s10745-017-9927-0
- Mukasa-Mugerwa, E, Lahlou-Kassi, A, Anindo, D, Rege, JE, Tembely, S, Tibbo, M and Baker, R. 2000. ‘Between and within breed variation in lamb survival and the risk factors associated with major causes of mortality in indigenous Horro and Menz sheep in Ethiopia’. Small Rumin. Res., 37: 1–12. DOI: 10.1016/S0921-4488(99)00152-2
- Munson, PJ. 2000. ‘Age-correlated Differential Destruction of Bones and its Effect on Archaeological Mortality Profiles of Domestic Sheep and Goats’. J. Archaeol. Sci., 27: 391–407. DOI: 10.1006/jasc.1999.0463
- Munson, PJ and Garniewicz, RC. 2003. ‘Age-mediated Survivorship of Ungulate Mandibles and Teeth in Canid-ravaged Faunal Assemblages’. J. Archaeol. Sci., 30: 405–416. DOI: 10.1006/jasc.2002.0850
- Negassa, A, Gebremedhin, B, Desta, S, Nigussie, K, Gebru, G, Shapiro, B, Dutilly-Diane, C and Tegegne, A. 2015.
‘Integrated bio-economic simulation model for goat production: Ex-ante evaluation of investment opportunities in Ethiopia’ . Nairobi, Kenya Int. Livest. Res. Inst. LIVES Work. - Nitsiakos, V. 1985. A Vlach pastoral community in Greece. University of Cambridge.
- Payne, S. 1973. ‘Kill-off Patterns in Sheep and Goats: The Mandibles from Aşvan Kale’. Anatol. Stud., 23: 281–303. DOI: 10.2307/3642547
- Price, M, Wolfhagen, J and Otárola-Castillo, E. 2016. ‘Confidence Intervals in the Analysis of Mortality and Survivorship Curves in Zooarchaeology’. Am. Antiq., 81: 157–173. DOI: 10.7183/0002-7316.81.1.157
- R Core Team. 2025. R: A Language and Environment for Statistical Computing.
- Redding, RW. 1981.
Decision making in subsistence herding of sheep and goats in the Middle East , University of Michigan. - Redding, RW. 1984.
‘Theoretical determinants of a herder’s decisions: Modeling variation in the sheep/goat ratio’ . In: Clutton-Brock, J and Grigson, C (eds.) Animals and Archaeology 3, Early Herders and Their Flocks, British Archaeological Reports 202, pp. 223–242. - Rosa, HJD and Bryant, MJ. 2002. ‘The ‘ram effect’ as a way of modifying the reproductive activity in the ewe’. Small Rumin. Res., 45: 1–16. DOI: 10.1016/S0921-4488(02)00107-4
- Sierra, A, Balasse, M, Radović, S, Orton, D, Fiorillo, D and Presslee, S. 2023. ‘Early Dalmatian farmers specialized in sheep husbandry’. Sci. Rep., 13:
10355 . DOI: 10.1038/s41598-023-37516-z - Singh, MK, Rai, B and Sharma, N. 2011. ‘Factors affecting survivability of Jamunapari kids under semi-intensive management system’. Indian J. Anim. Sci., 78.
- Sobol’, IM. 1990. ‘On sensitivity estimation for nonlinear mathematical models’. Math. Model., 112–118.
- Talore, D, Abebe, G and Tegegne, A. 2014. ‘Non-genetic factors influencing reproductive traits and preweaning mortality of lambs and kids under smallholder management, Southern Ethiopia’. J. Anim. Plant Sci., 24: 413–417.
- Tornero, C, Balasse, M, Bréhard, S, Carrère, I, Fiorillo, D, Guilaine, J, Vigne, J-D and Manen, C. 2020. ‘Early evidence of sheep lambing de-seasoning in the Western Mediterranean in the sixth millennium BCE’. Sci. Rep., 10:
12798 . DOI: 10.1038/s41598-020-69576-w - Triozzi, NP. 2024. Herding is Risky Business: Agropastoral Strategies in Neolithic Northern Dalmatia. University of California, Santa Barbara.
- Triozzi, NP. 2025. Age-at-Death Analysis of Caprine mandibles from Neolithic Sites in Dalmatia [Data set]. Zenodo. DOI: 10.5281/zenodo.17697101
- Upton, M. 1984.
‘Models of improved production systems for small ruminants’ . In: Sumberg, JE and Cassaday, K (eds.) Proceedings of the Workshop on Small Ruminant Production Systems in the Humid Zone of West Africa. International Livestock Centre for Africa, Ibadan, Nigeria. - Vigne, J-D and Helmer, D. 2007. ‘Was milk a “secondary product” in the Old World Neolithisation process? Its role in the domestication of cattle, sheep and goats’. Anthropozoologica, 42: 9–40.
- Watson, RH and Radford, HM. 1960. ‘The influence of rams on onset of oestrus in Merino ewes in the spring’. Aust. J. Agric. Res., 11: 65–71. DOI: 10.1071/AR9600065
- Wilson, RT. 1978. ‘Livestock Production on Masai Group Ranches’. East African Agric. For. J., 43: 193–199. DOI: 10.1080/00128325.1978.11662900
- Wilson, RT. 1989. ‘Reproductive performance of African indigenous small ruminants under various management systems: a review’. Anim. Reprod. Sci., 20: 265–286. DOI: 10.1016/0378-4320(89)90075-4
- Wilson, RT and Durkin, JW. 1983. ‘Livestock production in central Mali: weight at first conception and ages at first and second parturitions in traditionally managed goats and sheep’. J. Agric. Sci., 100: 625–628. DOI: 10.1017/S0021859600035383
- Wilson, RT, Peacock, C and Sayers, AR. 1984. ‘Aspects of reproduction in goats and sheep in south-central Kenya’. Anim. Sci., 38: 463–467. DOI: 10.1017/S0003356100041660
- Winterhalder, B, Lu, F and Tucker, B. 1999. ‘Risk-Sensitive Adaptive Tactics: Models and Evidence from Subsistence Studies in Biology and Anthropology’. J. Archaeol. Res., 7: 301–348. DOI: 10.1007/BF02446047
- Zeder, M. 2006. ‘Reconciling rates of long bone fusion and tooth eruption and wear in sheep (Ovis) and goat (capra)’. In: Ruscillo, D (ed.) Recent Advances in Ageing and Sexing Animal Bones. Proceedings of the 9th ICAZ Conference, Durham 2002, pp. 87–118. DOI: 10.2307/j.ctvh1ds02.10
- Zeder, MA and Pilaar, SE. 2010. ‘Assessing the reliability of criteria used to identify mandibles and mandibular teeth in sheep, Ovis, and goats, Capra’. J. Archaeol. Sci., 37: 225–242. DOI: 10.1016/j.jas.2009.10.002
DOI: https://doi.org/10.5334/jcaa.257 | Journal eISSN: 2514-8362
Language: English
Page range: 225 - 244
Submitted on: Nov 26, 2025
Accepted on: Apr 21, 2026
Published on: Jun 10, 2026
Published by: Ubiquity Press
In partnership with: Paradigm Publishing Services
Publication frequency: 1 issue per year
© 2026 Nicholas Triozzi, Sarah B. McClure, published by Ubiquity Press
This work is licensed under the Creative Commons Attribution 4.0 License.