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Changing Movements in a Changing World: Modelling Early Pleistocene and Early Middle Pleistocene Climatic and Ecological Environments and Influences on Hominin Dispersal in Eurasia Cover

Changing Movements in a Changing World: Modelling Early Pleistocene and Early Middle Pleistocene Climatic and Ecological Environments and Influences on Hominin Dispersal in Eurasia

Open Access
|Feb 2026

Figures & Tables

Table 1

Overview of the climatic models presented in this paper, emphasising the parameters, spatial and temporal resolution and time reach of the models. The column to the far right shows the sources of information for the model.

CLIMATIC PARAMETERSSPATIAL RESOLUTION (LATITUDE × LONGITUDE)TEMPORAL RESOLUTIONPERIOD REACHSOURCEAVAILABLE SITE
CESM2Atmosphere, ocean, land, river run-off, land ice, sea ice1° × 1°
or 2° × 2°
1 kyrBacmeister et al. (2020),
Rocha (2023),
Cesm.ucar.edu
Community Earth System Model 2 (CESM2) | Community Earth System Model
HadCM3Atmosphere and ocean temperature,
sea ice quantity
2.5° × 3.75°1 kyr800 kaGordon et al. (2000),
Krapp et al. (2021)
BRIDGE — Paper: Valdes et al 2017
Paleo-PGEMOcean-atmospheric model (AOGCM) & ice sheet quantity1° × 1°1 kyr5 MaHolden et al. (2016, 2019),
Barreto et al. (2023)
PALEO-PGEM-Series: a spatial time series of the global climate over the last 5 million years (Plio-Pleistocene)
Paleo-ClimSurface temperature & precipitation estimates (AOGCM)0.04° × 0.04°Snapshots130 ka, 787 ka, 3.264–3.025 Ma, 3.3 MaBrown et al. (2018)PaleoClim.org
Oscillayers19 bio-climatic parameters of mean and extreme temperatures and precipitation0.04° × 0.04°10 kyr5.4 MaGamisch (2019)Dryad | Data -- Oscillayers: a data set for the study of climatic oscillations over Plio-Pleistocene time scales at high spatial-temporal resolution
PMIP4Air temperature, precipitation, sea surface temperature, ocean heat, snow cover & depth, soil moisture, ice sheet extent, vegetation fraction0.5° × 0.5° to 2.5° × 2.5°SnapshotsLast millennium, LGM, mid-Holocene, last glacial, mid-PlioceneTaylor et al. (2012), Kageyama et al. (2018)Model database | PMIP
Table 2

Overview of the environmental reconstructions presented in this paper, emphasising the parameters, spatial and temporal resolution, and chronological range of the models. The column to the far right shows the sources used for the reconstructions.

CLIMATIC PARAMETERSOUTPUTSSPATIAL RESOLUTIONTEMPORAL RESOLUTIONPERIOD REACHSOURCEAVAILABLE SITE
LOVECLIM/iLOVECLIMOcean general circulation model (AOGCM) & thermodynamic sea ice modelVegetation cover following climatic variables1° × 1°1-year125 kaGoosse et al. (2010), Roche et al. (2014)Loveclim 784K | Climate Data
Overview – iLOVECLIM ESM
PRISM4Palaeogeography, sea level, ocean temperature, land and sea iceTerrestrial vegetation, soil, lakes0.25° × 0.25°Marine isotope stages2.6–3.6 MaDowsett et al. 2016PRISM4: Data
BIOME4Atmospheric CO2 (CESM)Vegetation cover, divided into 28 biomes from PFTs0.5° × 0.5°1000-yearLGMHarrison & Prentice (2003),
Zeller et al. (2023)
GitHub – jedokaplan/BIOME4: The BIOME4 equilibrium global vegetation model
CARAIBAir temperature, precipitation, sunshine hours, air humidity, wind speedSoil hydrology, surface energy budget, GPP, NPP, LAI, biomass, BAGs cover fraction, fire module1° × 1°1-yearLGM, Mid Holocene, MioceneFrançois et al. (1998), Dury et al. (2011)UMCCB: Research: Projects: BIOSERF
ORCHIDEEAtmospheric CO2, air temperature, humiditySoil carbon, soil temperature and hydrology, vegetation divided into 13 PFT groups, river and floodplain scheme1° × 1°1-yearInput dependentKrinner et al. (2005, Guimberteau et al. (2018)Documentation/Forcings – ORCHIDEE
jcaa-9-1-212-g1.png
Figure 1

Flow diagram displaying the workflow for producing a raster layer applicable to a hominin dispersal simulation.

Table 3

List of climatic emulators and reconstructions addressed in this paper, advantages and limitations for investigating global hominin dispersal patterns during the Early Pleistocene and validation methods.

EMULATORADVANTAGESLIMITATIONSVALIDATION
CESM2High spatial resolution
High temporal resolution
Lack of Pleistocene studies/dataNo, but comes with control simulations
HadCM3High temporal resolutionLower spatial resolution
Timescale limited to 800 ka
Observed ocean data
Paleo-PGEMHigh spatial resolution
High temporal resolution
Time reach: 5 Ma
Computing limitations, accuracy of represented elements/dynamics in the downscalingObserved temperature reconstructions & proxy data
Paleo-ClimVery high spatial resolutionSnapshot approachInter-model comparison
OscillayersVery high spatial resolution
Bioclimatic parameters
Time reach: 5.4 Ma
Low temporal resolutionInter-model comparison
PMIP4Very high spatial resolution
Detailed output
Snapshot approach
Requires detailed input data
Proxy reconstructions, control simulations, benchmark tools
Table 4

List of environmental emulators and reconstructions addressed in this paper, advantages and limitations for investigating global hominin dispersal patterns during the Early Pleistocene and validation methods.

EMULATORADVANTAGESLIMITATIONSVALIDATION
LOVECLIM/
iLOVECLIM
High spatial resolution
High temporal resolution
Vegetation cover
Sea Ice
So far only tested back to 125 kaProxy data & inter-model comparison
PRISM4Very high spatial resolution
Vegetation cover
Land and sea ice
Lake element
Time reach: 2.6–3.6 Ma
Low temporal resolutionNo, functions as a conceptual model
BIOME4Very high spatial resolution
High temporal resolution
Biome-divided vegetation cover
Limited to the LGMProxy data divided into PFT groups
CARAIBVery high spatial resolution
Variety of output data: soil hydrology, biomass, GPP, NPP, LAI & BAGs
Input data requirementsInter-model comparisons, CO2 observations
ORCHIDEEHigh spatial resolution
Very high temporal resolution
PFT-class divided vegetation cover, including GPP and NPP values
River and floodplain scheme
Applicable to any climate forcing
Very detailed, heavy to run
Based on present measurements
Observed temperature gradients, basin-scale averages, topsoil moisture models, forest data
DOI: https://doi.org/10.5334/jcaa.212 | Journal eISSN: 2514-8362
Language: English
Submitted on: Mar 5, 2025
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Accepted on: Sep 5, 2025
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Published on: Feb 27, 2026
Published by: Ubiquity Press
In partnership with: Paradigm Publishing Services
Publication frequency: 1 issue per year

© 2026 Kamilla L. Lomborg, Carolina Cucart-Mora, Jan-Olaf Reschke, Christine Hertler, Matt Grove, Benoit Gaudou, Mehdi Saqalli, Marie-Hélène Moncel, published by Ubiquity Press
This work is licensed under the Creative Commons Attribution 4.0 License.