References
- Abbott, J. C., & Dill, L. M. (1985). Patterns of aggressive attack in juvenile steelhead trout (Salmo gairdneri). Canadian Journal of Fisheries and Aquatic Sciences, 42(11), 1702–1706. https://doi.org/10.1139/f85-213
- Agnew, D. J. (1988). Evidence for the existence of two populations of Irish Sea cod (Gadus morhua L.) from consideration of growth rates. ICES Committee Meeting Papers 1988/G, 65, 20. https://doi.org/10.15835/nsb.8.4.9852
- Al Ramahi, F. K., & Al Bahadly, Z. K. I. (2017). Estimation of Suaeda aegyptiaca plant distribution regions at Iraq using RS & GIS applications. Iraqi Journal of Science, 58(2A), 767–777.
- Aleev, Y. G. (1969). Function and gross morphology in fish (p. 268). Israel Program for Scientific Translations.
- Al-Faisal, A. J., & Mutlak, F. M. (2015). First record of the Nile tilapia Oreochromis niloticus (Linnaeus, 1758), from the Shatt Al-Arab River, Southern Iraq. International Journal of Marine Science, 5, 1–3. https://doi.org/10.5376/ijms.2015.05.0038
- Ali, F. R., Fouad, K. M., & Abdul-Rahman, B. A. (2019). Evaluation of impact of vegetation decrease on precipitation rates in Baghdad CITY using remote sensing technique. Ecology, Environment and Conservation, 25, S44–S50.
- Allendorf, F. W. (1988). Conservation biology of fishes. Conservation Biology, 2(2), 145–148. https://doi.org/10. 1111/j.1523-1739.1988.tb00165.x
- Al-Zaidy, K. J. (2013). First record of Tilapia zilli (Gewais, 1848) in Al-Delmj Marsh west Al-Diwania city middle of Iraq. Diyala Agricultural Sciences Journal, 5(1), 9–16.
- Anane-Taabeah, G. (2019). Characterization of the molecular genetic variation in wild and farmed Nile tilapia Oreochromis niloticus in Ghana for conservation and aquaculture development. Dissertation submitted to the faculty of the Virginia Polytechnic Institute and State University in partial fulfilment of the requirements for the degree of Doctor of Philosophy in Fisheries Sciences (Department of Fish and Wildlife Conservation) (p. 115). The Virginia Polytechnic Institute and State University.
- Avsar, D. (1994). A stock differentiation study of the sprat (Sprattus sprattus phalericus Risso) off the southern coast of the Black Sea. Fisheries Research, 19(3–4), 363–378. https://doi.org/10.1016/0165-7836(94)90050-7
- Barriga-Sosa, I. D. L. A., Jiménez-Badillo, M. D. L., Ibanez, A. L., & Arredondo-Figueroa, J. L. (2004). Variability of tilapias (Oreochromis spp.) introduced in Mexico: Morphometric, meristic and genetic characters. Journal of Applied Ichthyology, 20(1), 7–14. https://doi.org/10.1111/j.1439-0426.2004.00445.x
- Beecham, R. V., Minchew, C. D., & Parsons, G. R. (2007). Comparative swimming performance of juvenile pond-cultured and wild-caught channel catfish. North American Journal of Fisheries Management, 27(3), 729–734. https://doi.org/10.1577/M04-219.1
- Boglione, C., Costa, C., Di Dato, P., Ferzini, G., Scardi, M., & Cataudella, S. (2003). Skeletal quality assessment of reared and wild sharp snout sea bream and pandora juveniles. Aquaculture, 227(1–4), 373–394. https://doi.org/10.1016/S0044-8486(03)00515-5
- Booke, H. E. (1981). The conundrum of the stock concept—Are nature and nurture definable in fishery science? In R. C. Simon, & P. A. Larkin (Eds), The stock concept in Canadian fisheries (pp. 11–14). Canadian Special Publication of Fisheries and Aquatic Sciences, 59. https://doi.org/10.1139/f81-200
- Bosakowski, T., & Wagner, E. J. (1994). Assessment of fin erosion by comparison of relative fin length in hatchery and wild trout in Utah. Canadian Journal of Fisheries and Aquatic Sciences, 51(3), 636–641. https://doi.org/10.1139/f94-064
- Bostanci, D., Yedier, S., Helli, S., & Polat, N. (2021). Ordu ili iç sularında Carassius auratus (Linnaeus, 1758) türünün ilk kaydı ve türün Ulugöl Yaylası Göleti popülasyonuyla ilgili bazı veriler. Aquatic Research, 4(3), 279–285. https://doi.org/10.3153/AR21021
- Brönmark, C., & Pettersson, L. B. (1994). Chemical cues from piscivores induce a change in morphology in crucian carp. Oikos, 70(3), 396–402. https://doi.org/10.2307/3545777
- Bronte, C. R., Fleischer, G.W., Maistrenko, S. G., & Pronin, N. M. (1999). Stock structure of Lake Baikal omul as determined by whole body morphology. Journal of Fish Biology, 54(4), 787–798. https://doi.org/10.1111/j.1095-8649.1999.tb02033.x
- Coad, B. W. (1996). Exotic and transplanted fishes in Southwest Asia. Proceedings of the 8th Congress of Societas Europaea Ichthyologorum (SEI) (pp. 81–106). 26 September - 26–22 October 1994, 1996.
- Corti, M., Thorpe, R. S., Sola, L., Sbordoni, V., & Cataudella, S. (1988). Multivariate morphometrics in aquaculture: A case study for six stocks of the common carp (Cyprinus carpio) from Italy. Canadian Journal of Fisheries and Aquatic Sciences, 45, 1548–1554. https://doi.org/10.1139/f88-183
- Creech, S. (1992). A multivariate morphometric investigation of Atherina boyeri Risso, 1810 and A. presbyter Cuvier, 1829 (Teleostei: Atherinidae): Morphometric evidence in support of the two species. Journal of Fish Biology, 41(3), 341–353. https://doi.org/10.1111/j.1095-8649.1992.tb02663.x
- Cruz, E. M., & Ridha, M. (1994). Overwintering tilapia, Oreochromis spilurus (Günther), fingerlings using warm underground sea water. Aquaculture Research, 25(9), 865–871. https://doi.org/10.1111/j.1365-2109.1994.tb01348.x
- Cuadras, C. M. (2019). Nuevos Métodos de Análisis Multivariante (pp. 89–90). CMC Editions. (in Spanish).
- Currens, K. P., Sharpe, C. S., Hjort, R., Schreck, C. B., & Li, H. W. (1989). Effects of different feeding regimes on the morphometrics of chinook salmon (Oncorhynchus tshawytscha) and rainbow trout (O. mykiss). Copeia, 3(3), 689–695. https://doi.org/10.2307/1445496
- Davidson, K., Roff, J. C., & Elner, R. W. (1985). Morphological, electrophoretic and fecundity characteristics of Atlantic snowcrab, Chionoecetes opilio, and implications for fisheries management. Canadian Journal of Fisheries and Aquatic Sciences, 42(3), 474–482. https://doi.org/10.1139/f85-064
- Doyle, R. W., Shackel, N. L., Basiao, Z., Uraiwan, S., Matricia, T., & Talbot, A. J. (1991). Selective diversification of aquaculture stocks: A proposal for economically sustainable genetic conservation. Canadian Journal of Fisheries and Aquatic Sciences, 48(Suppl. 1), 148–154. https://doi.org/10.1139/f91-313
- Einum, S., & Fleming, I. A. (1997). Genetic divergence and interactions in the wild among native, farmed and hybrid Atlantic salmon. Journal of Fish Biology, 50(3), 634–651. https://doi.org/10.1111/j.1095-8649.1997.tb01955.x
- Elliott, J. M. (1990). Mechanisms responsible for population regulation in young migratory trout, Salmo trutta. III. The role of territorial behaviour. The Journal of Animal Ecology, 59(3), 803–818. https://doi.org/10.2307/5015
- El-Serafy, S. S., Abdel-Hameid, N. A. H., Awwad, M. H., & Azab, M. S. (2007). DNA riboprinting analysis of tilapia species and their hybrids using restriction fragment length polymorphisms of the small subunit ribosomal DNA. Aquaculture Research, 38(3), 295–303. https://doi.org/10.1111/j.1365-2109.2007.01668.x
- El-Zaeem, S. Y., Ahmed, M. M. M., Salama, M. E. S., & El-Kader, W. N. A. (2012). Phylogenetic differentiation of wild and cultured Nile tilapia (Oreochromis niloticus) populations based on phenotype and genotype analysis. African Journal of Agricultural Research, 7(19), 2946–2954. https://doi.org/10.5897/AJAR11.2170
- Fagen, R., & Smoker, W. W. (1989). How large-capacity hatcheries can alter interannual variability of salmon production. Fisheries Research, 8(1), 1–11. https://doi.org/10.1016/0165-7836(89)90036-2
- FAO. (2023). The ELEFAN software for length-frequency analysis in R: Tutorial and manual. FAO Fisheries and Fisheries Aquaculture Technical Paper No. 682. FAO. https://doi.org/10.4060/cc3223en
- Fleming, I. A., & Einum, S. (1997). Experimental tests of genetic divergence of farmed from wild Atlantic salmon due to domestication. ICES Journal of Marine Science, 54(6), 1051–1063. https://doi.org/10.1016/S1054-3139(97)80009-4
- Fleming, I. A., Jonsson, B., & Gross, M. R. (1994). Phenotypic divergence of sea-ranched, farmed, and wild salmon. Canadian Journal of Fisheries and Aquatic Sciences, 51(12), 2808–2824. https://doi.org/10.1139/f94-280
- Foote, C. J., Moore, K., Stenberg, K., Craig, K. J., Wenburg, J. K., & Wood, C. C. (1999). Genetic differentiation in gill raker number and length in sympatric anadromous and nonanadromous morphs of sockeye salmon, Oncorhynchus nerka. Environmental Biology of Fishes, 54(3), 263–274. https://doi.org/10.1023/A:1007548807233
- Galemoni de Graaf, G. J. F., & Huisman, E. A. (1999). Reproductive biology of pond reared Nile tilapia, Oreochromis niloticus L. Aquaculture Research, 30(1), 25–33. https://doi.org/10.1046/j.1365-2109.1999.00295.x
- Gjedrem, T., Gjerde, B., & Refstie, T. (1988). A review of quantitative genetic research in salmonids at AKVAFORSK. In B. S. Weir, E. Eisen Jr., M. M. Good’man, & G. Namkoong (Eds.), Proceedings of the second international conference on quantitative genetics (pp. 527–535). Sinauer.
- Gjerde, B. A., & Schaeffer, L. R. (1989). Body traits in rainbow trout: II. Estimates of heritabilities and of phenotypic and genetic correlations. Aquaculture, 80(1–2), 25–44. https://doi.org/10.1016/0044-8486(89)90271-8
- Gupta, M. V., & Acosta, B. O. (2004). A review of global tilapia farming practices. Aquaculture Asia, 9(1), 7–12.
- Hanson, K. C., Hasler, C. T., Suski, C. D., & Cooke, S. J. (2007). Morphological correlates of swimming activity in wild largemouth bass (Micropterus salmoides) in their natural environment. Comparative Biochemistry and Physiology A, 148(4), 913–920. https://doi.org/10.1016/j. cbpa.2007.09.013
- Hassanien, H. A., Kamel, E. A., Salem, M. A., & Dorgham, A. S. (2011). Multivariate analysis of morphometric parameters in wild and cultured Nile tilapia Oreochromis niloticus. Journal of the Arabian Aquaculture Society, 6(2), 424–440.
- Hedgecock, D., Hutchinson, E. S., Li, G., Sly, F. L., & Nelson, K. (1989). Genetic and morphometric variation in the Pacific sardine, Sardinops sagax caerulea: Comparisons and contrasts with historical data and with variability in the northern anchovy, Engraulis mordax. Fishery Bulletin, 87(3), 653–671.
- Hockaday, S., Beddow, T. A., Stone, M., Hancock, P., & Ross, L. G. (2000). Using truss networks to estimate the biomass of Oreochromis niloticus, and to investigate shape characteristics. Journal of Fish Biology, 57(4), 981–1000. https://doi.org/10.1111/j.1095-8649.2000.tb02206.x
- Hubbs, C. L., & Lagler, K. F. (1958). Fishes of the great lakes region. University of Michigan Press.
- Ikpeme, E. V., Ekerette, E. E., Udensi, O. U., & Ozoje, M. O. (2017). Assessment of morphological variation in wild and cultured populations of tilapia Fish (Oreochromis niloticus). Journal of Advances in Biology and Biotechnology, 13(2), 1–10. https://doi.org/10.9734/JABB/2017/33777
- Jawad, L. A., Ibáñez, A. L., Kiki, M., & Gnohossou, P. (2020). Determination of body shape and meristic characters variations in wild and cultured populations of cichlid fish, Oreochromis niloticus, from the Republic of Benin, West of Africa. Fisheries & Aquatic Life, 28(3), 186–194. https://doi.org/10.2478/aopf-2020-0022
- Johnsson, J. I., Petersson, E., Jönsson, E., Björnsson, B. T., & Järvi, T. (1996). Domestication and growth hormone alter antipredator behaviour and growth patterns in juvenile brown trout, Salmo trutta. Canadian Journal of Fisheries and Aquatic Sciences, 53(7), 1546–1554. https://doi.org/10.1139/f96-090
- Karaiskou, N., Triantafyllidis, A., Katsares, V., Abatzopoulos, T. J., & Triantaphyllidis, C. (2009). Microsatellite variability of wild and farmed populations of Sparus aurata. Journal of Fish Biology, 74(8), 1816–1825. https://doi.org/10.1111/j.1095-8649.2009.02186.x
- Kinsey, S. T., Orsoy, T., Bert, T. M., & Mahmoudi, B. (1994). Population structure of the Spanish sardine Sardinella aurita: Natural morphological variation in a genetically homogeneous population. Marine Biology, 118(2), 309–317. https://doi.org/10.1007/BF00349798
- Lehoczky, I., Al Fatle, F. A., Nguyen Thi, Q., Edviné Meleg, E., Sallai, Z., Szabó, G., Fekete, G., Kópor, I., Várkonyi, E., Péter, D., & Beliczky, G. (2025). Genetic survey of crucian carp Carassius carassius populations in Hungary for a conservation project to establish live gene bank. Scientific Reports, 15(1), 8835. https://doi.org/10.1038/s41598-025-93468-6
- Lindsey, C. C. (1981). Stocks are chameleons: Plasticity in gill rakers of coregonid fishes. Canadian Journal of Fisheries and Aquatic Sciences, 38(12), 1497–1506. https://doi.org/10.1139/f81-202
- Mamuris, Z., Apostolidis, A. P., Panagiotaki, P., Theodorou, A. J., & Triantaphllidis, C. (1998). Morphological variation between red mullet populations in Greece. Journal of Fish Biology, 52, 107–117. https://doi.org/10.1111/j.1095-8649.1998.tb01556.x
- Matsuoka, M. (1987). Development of skeletal tissue and skeletal muscle in the Red Sea bream, Pagrus major. Bulletin of the Seikai Regional Fisheries Research Laboratory, 65(3), 1–114.
- Matthews, W. J., & Robison, H. W. (1988). The distribution of the fishes of Arkansas: A multivariate analysis. Copeia, 2, 358–374. https://doi.org/10.2307/1445876
- Metcalfe, N. B., Huntingford, F. A., & Thorpe, J. E. (1988). Feeding intensity, growth rates, and the establishment of life-history patterns in juvenile Atlantic salmon Salmo salar. Journal of Animal Ecology, 57(2), 463–474. https://doi.org/10.2307/4918
- Muslih, K. D., & Abbas, A. M. (2024). Climate of Iraq. The geography of Iraq Salih Muhammad Awadh, Moutaz Al-Dabbas (ed.). (pp. 19–47). Springer Nature Switzerland. https://doi.org/10.1007/978-3-031-71356-9_2
- Mutlak, F. M., & Al-Faisal, A. J. (2009). A new record of two exotic cichlids fish Oreochromis aureus Steindacher, 1864 and Tilapia zillii Gervais, 1848 from south of the main outfall drain in Basrah city. Mesopotamian Journal of Marine Science, 24(2), 160–170.
- Narváez, J. C., Acero, A., & Blanco, J. (2005). Morphometric variation in naturalized and domesticated populations of the Nile tilapia Oreochromis niloticus (Teleostei: Cichlidae) in Northern Colombia. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales, 29(112), 383–394. (in Spanish). https://doi.org/10.18257/raccefyn.29(112)0.2005.2172
- Ndiwa, T. C., Nyingi, D. W., Claude, J., & Agnèse, J. F. (2016). Morphological variations of wild populations of Nile tilapia (Oreochromis niloticus) living in extreme environmental conditions in the Kenyan Rift-Valley. Environmental Biology of Fishes, 99(4), 473–485. https://doi.org/10.1007/s10641-016-0492-y
- Pakkasmaa, S. (2000). Morphological and early life history variation in salmonid fishes. Ph.D. Dissertation, Department of Ecology and Systematics. University of Helsinki.
- Parish, B. B., & Sharman, D. P. (1958). Some remark on method used in herring ‘racial’ investigations, with special reference to otolith studies. Rapports et procès-verbaux des réunions, International Council for the Exploration of the Sea, 143(1), 66–80.
- Penña-Mendoza, B., Gómez-Márquez, J. L., Salgado-Ugerte, I. H., & Ramírez-Nogguera, D. (2005). Reproductive biology of Oreochromis niloticus (Perciformes: Cichlidae) at Emiliano Zapata dam, Morelos, Mexico. Revista de Biologia Tropical, 53(3–4), 515–522. https://doi.org/10.15517/rbt.v53i3-4.14666
- Plaut, I. (2001). Critical swimming speed: Its ecological relevance. Comparative Biochemistry and Physiology A, 131(1), 41–50. https://doi.org/10.1016/S1095-6433(01)00462-7
- Ryman, N., Lagercrantz, U., Andersson, L., Chakraborty, R., & Rosenberg, R. (1984). Lack of correspondence between genetic and morphologic variability patterns in Atlantic herring (Clupea harengus). Heredity, 53(3), 687–704. https://doi.org/10.1038/hdy.1984.127
- Ryman, N., Utter, F., & Hindar, K. (1995). Introgression, supportive breeding, and genetic conservation. In J. D. Ballou, M. Gilpin, & T. J. Foose (Eds.), Population management for survival and recovery. Analytical methods and strategies in small population conservation (pp. 341–365). Columbia University Press.
- Saleh, K. I. (2007). First recorded of Tilapia zillii (Gervais, 1848), in natural water of Iraq (Tigris River) (pp. 26–27). The First Scientific Conference of Agricultures College, University of Basra.
- Sharp, J. J., Sargent, E. E., & Schweitzer, P. A. (2002). Genetic monitoring. In J. G. Fox, L. C. Anderson, F. M. Loew, & F. W. Quimby (Eds.), Laboratory Animal Medicine (2nd ed., pp. 1117–1127). Academic Press. https://doi.org/10.1016/B978-012263951-7/50030-2
- Shepherd, G. (1991). Meristic and morphometric variation in Black Sea bass north of Cape Hatteras, North Carolina. North American Journal of Fisheries Management, 11(2), 139–148. https://doi.org/10.1577/1548-8675(1991)011>;0139:MAMVIB>2.3.CO;2
- Solomon, S. G., Okomoda, V. T., & Ogbenyikwu, A. I. (2015). Intraspecific morphological variation between cultured and wild Clarias gariepinus (Burchell) (Clariidae, Siluriformes). Fisheries and Aquatic Life, 23(1), 53–61. https://doi.org/10.1515/aopf-2015-0006
- Stearns, S. C. (1989). The evolutionary significance of phenotypic plasticity. Bioscience, 39(7), 436–445. https://doi.org/10.2307/1311135
- Swain, D. P., & Foote, C. J. (1999). Stocks and chameleons: The use of phenotypic variation in stock identification. Fisheries Research, 43(1–3), 113–128. https://doi.org/10.1016/S0165-7836(99)00069-7
- Swain, D. P., Riddell, B. E., & Murray, C. B. (1991). Morphological differences between hatchery and wild populations of coho salmon (Oncorhynchus kisutch): Environmental versus genetic origin. Canadian Journal of Fisheries and Aquatic Sciences, 48(9), 1783–1791. https://doi.org/10.1139/f91-210
- Taylor, E. B. (1991). A review of local adaptation in salmonidae, with particular reference to Pacific and Atlantic salmon. Aquaculture, 98(1–3), 185–207. https://doi.org/10.1016/0044-8486(91)90383-I
- Teugels, G. G., & Vreven, E. (1997). Preliminary results on morphometric differentiation between natural populations of the Nile tilapia Oreochromis niloticus (Perciformes, Cichlidae). Proceedings of the Symposium Genetics and Aquaculture in Africa, Abidjan (pp. 1–4).
- Thompson, J. D. (1991). Phenotypic plasticity as a component of evolutionary change. Trends in Ecology and Evolution, 6(8), 246–249. https://doi.org/10.1016/0169-5347(91)90070-E
- Thorpe, J. E. (1986). Age at first maturity in Atlantic salmon, Salmo salar: Freshwater period influences and conflicts with smolting. In D.J. Meerburg (Ed.), Salmonid age at maturity. Canadian Special Publication of Fisheries and Aquatic Sciences, 89, 7–14.
- Tibihika, P. D., Curto, M., Alemayehu, E., Waidbacher, H., Masembe, C., Akoll, P., & Meimberg, H. (2020). Molecular genetic diversity and differentiation of Nile tilapia (Oreochromis niloticus, L. 1758) in East African natural and stocked populations. BMC Evolutionary Biology, 20(1), 16. https://doi.org/10.1186/s12862-020-1583-0
- Turan, C., Erguden, D., Turan, F., & Gurlek, M. (2004). Genetic and morphologic structure of Liza abu (Heckel, 1843) populations from the rivers Orontes, Euphrates and Tigris. Turkish Journal of Veterinary and Animal Sciences, 28(4), 729–734.
- Turan, C., Oral, M., Ozturk, B., & Duzgunes, E. (2006). Morphometric and meristic variation between stocks of bluefish (Pomatomus saltatrix) in the Black, Marmara, Aegean and Northeastern Mediterranean Seas. Fisheries Research, 79(1-2), 139–147. https://doi.org/10.1016/j.fishres.2006.01.015
- Vreven, E. M., Adepo-Gourene, B., Agnèse, J. F., & Teugels, G. G. (1998). Morphometric and allozyme variation in natural populations and cultured strains of the Nile tilapia Oreochrornis niloticus (Teleostei, Cichlidae). Belgian Journal of Zoology, 128(1), 23–34.
- Wainwright, P. C., Osenberg, C. W., & Mittelbach, G. G. (1991). Trophic polymorphism in the pumpkinseed sunfish (Lepomis gibbosus Linnaeus): Effects of environment on ontogeny. Functional Ecology, 5(1), 40–55. https://doi.org/10.2307/2389554
- West-Eberhard, M. J. (1989). Phenotypic plasticity and the origins of diversity. Annual Review of Ecology, Evolution, and Systematics, 20(1), 249–278. https://doi.org/10.1146/annurev.es.20.110189.001341
- Wimberger, P. H. (1992). Plasticity of fish body shape. The effects of diet, development, family and age in two species of Geophagus (Pisces: Cichlidae). Biological Journal of the Linnean Society, 45(3), 197–218. https://doi.org/10.1111/j.1095-8312.1992.tb00640.x https://www.academia.edu/41705592/Nuevos_m%C3%A9todos_de_an%C3%A1lisis_multivariante_Cuadras
- Yi, Y., Lin, C. K., & Diana, J. S. (1996). Influence of Nile tilapia (Oreochromis niloticus) stocking density in cages on their growth and yield in cages and ponds containing the cages. Aquaculture, 146(3–4), 205–215. https://doi.org/10.1016/S0044-8486(96)01377-4
- Youngson, A. F., Martin, S. A. M., Jordan, W. C., & Verspoor, E. (1991). Genetic protein variation in Atlantic salmon in Scotland: Comparison of wild and farmed fish. Aquaculture, 98(1–3), 231–242. https://doi.org/10.1016/0044-8486(91)90386-L