Figure 1

Figure 2

Figure 3

Figure 4

Chronological distribution data and ecological data from the literature and our sampling
| region | period | depth (m) | sediment | number | associates | source |
|---|---|---|---|---|---|---|
| Gold Coast (= Ghana) to Roscoff, including W Mediterranean Sea | 1932–1977 | 0–200 | mud, muddy sand, sandy mud | 1–40 | Cerianthus membranaceus (Gmelin, 1791), Sipunculus nudus Linnaeus, 1766, Oestergrenia digitata (Montagu, 1815) | Glémarec & Hily (1979), partly cited from earlier sources |
| Bay of the Seine | 2008–2011 | 10–25 | mud, muddy sand, sand | 30 | Terebellides stroemi Sars, 1835, Lanice conchilega Pallas, 1766, Upogebia deltaura (Leach, 1816) | Jourde et al. (2012) |
| Dieppe–Le Tréport | 2015–2018 | 12–25 | including coarse sand | Chaetopterus variopedatus (Renier, 1804), Cerianthus lloydii Gosse, 1859 | Pezy & Dauvin (2016) | |
| southern North Sea | 2019 | 32 | sand with shell grit | 1 | Callianassa subterranea and/or Gilvossius tyrrhenus, Echinocardium cordatum | this paper |