Fig. 1
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Fig. 8
Fig. 9
Measurements of Paraoxydirus vulvalpapillatus n_ sp_ (All the measurements in μm except L in mm)_
| Locality | Type population | Karnataka population | ||
|---|---|---|---|---|
| Characters | Holotype female | Paratype females | Paratype males | Female |
| n | 1 | 5 | 8 | 1 |
| L | 4.8 | 4.6± 0.3 (4.3 – 4.9) | 4.4±0.5 (3.8 – 5.0) (n=3) | 4.6 |
| Body width at neck base | 40 | 42±1.3 (41 – 44) | 42.1±2.4 (38 – 45) | 47 |
| Body width at mid body | 53 | 51.6±2.6 (47 – 55) | 48.6±4.7 (41 – 56) | 65 |
| Body width at anus/ cloaca | 31 | 28.6±2.4 (25 – 32) | 36±2.2 (32 – 39) | 32 |
| a | 92 | 88.9±4.9 (83.2 – 95.2) | 84.4± 10.7 (69.3 – 92.7) (n=3) | 71.6 |
| b | 14.3 | 14.1± 0.6 (13 – 15) | 14.1± 0.8 (13 – 15) (n=3) | 13.2 |
| c | 15 | 13.9±0.3 (13.5 – 14.3) | 14.2±5.0 (8.1 – 20.3) (n=3) | 14.9 |
| c’ | 10.5 | 11.2± 1.2 (9.7 – 12.6) | 9.8± 3.6 (6.8 – 14.8) (n=3) | 9.8 |
| V | 36.3 | 37.8± 1.6 (35.6 – 39.5) | - | 41.6 |
| G1 | 10.2 | 9.2± 1.9 (6.6 – 13.3) | - | 6.9 |
| G2 | 10.5 | 11.2± 0.7 (10.2 – 11.7) | - | 7.6 |
| Lip region width | 10 | 10.6±0.5 (10 – 11) | 10.3±0.5 (10 – 11) | 10 |
| Lip region height | 4 | 4.2±0.7 (3 – 5) | 4.5±0.5 (4 – 5) | 3 |
| Amphid aperture width | 5 | 5.0±0.6 (4 – 6 ) | 6.3±0.7 (5 – 7) | 5 |
| Odontostyle length | 5 | 5.2±0.4 (5 – 6) | 4.8±0.3 (4 – 5) | 5 |
| Odontophore length | 11 | 11.5±0.7 (11 – 13) | 12±0.7 (11 – 13) | 11 |
| Guiding ring from ant. End | 5 | 5.2±0.4 (5 – 6) | 5.3±0.5 (5 – 6) | 4 |
| Nerve ring from ant. End | 143 | 149.4±2.9 (145 – 153) | 145.1±6.1 (135 – 155) | 142 |
| Neck length | 340 | 331.8±11.6 (317 – 352) | 322.8±14.9 (297 – 339) | 352 |
| Expanded part of pharynx | 90 | 93±10.4 (82 – 107) | 86.3±10.6 (76 – 107) | 138 |
| Cardia length | 13 | 15.6±2.2 (12 – 19) | 13.1±2.5 (11 – 19) | 15 |
| Anterior genital branch | 499 | 469.2±102.6 (288 – 575) | - | 320 |
| Posterior genital branch | 512 | 514±33.2 (492 – 580) | - | 356 |
| Vaginal depth | 30 | 30.2±0.7 (29 – 31) | - | 34 |
| Vulva from ant. end | 1770 | 1703±115.8 (1550 – 1896) | - | 1934 |
| Prerectum length | 455 | 373.7±42.5 (315 – 435) | 486.7±61 (432 – 602) | 290 |
| Rectum length | 37 | 36.6±2.7 (32 – 40) | 46.7±2.6 (42 – 50) | 43 |
| Tail length | 325 | broken L = 310 – 365 | 341± 97(247 – 475) (n=3) (broken L= 185 – 279) | 312 |
| Spicule length | - | - | 51±2.5 (46 – 54) | - |
| Lateral guiding pieces | - | - | 12.5±1.6 (11 – 16) | - |
| Ventromedian supplements | - | - | 11 – 14 | - |
Measurements of Paraoxydirus novus Jairajpuri, 1965 (All the measurements in μm except L in mm)_
| Locality | Tamil Nadu population | Karnataka population | Goa population | Assam population |
|---|---|---|---|---|
| Characters | Females | Females | Females | Females |
| n | 6 | 4 | 5 | 5 |
| L | 3.9± 0.1 (3.7 - 4.0) (n=3) | 3.4 ± 0.1 (3.3 - 3.5) | 3.3±0.1 (3.2 - 3.5) (n=3) | 3.2± 0.1 (3.1 - 3.4) (n=3) |
| Body width at neck base | 52.6±3.8 (50 - 61) | 46.5±1.7 (44 - 48) | 49.4±3.8 (43 - 54) | 49.4± 4.3 (42 - 55) |
| Body width at mid body | 61.1±3.8 (57 - 69) | 57.7 ±3.4 (52 - 61) | 59.8±5.9 (49 - 67) | 55.4± 4.4 (50 - 62) |
| Body width at anus/ cloaca | 32.5±1.6 (31 - 36) | 31.7 ±1.1 (30 - 33) | 31.2±1.5 (30 - 33) | 29± 1.4 (27 - 31) |
| a | 61.7± 3.9 (56.7 - 66.2) (n=3) | 59 ±4.0 (55.6 - 65.8) | 55.3 ± 2.3 (52.4 - 58) (n=3) | 58.7 ± 5.3 (53 - 68.2) (n=3) |
| b | 11.7± 0.4 (11 - 12.2) (n=3) | 10.6±0.2 (10.2 - 10.8) | 10±0.4 (9.7 - 10.6) (n=3) | 10.6 ± 0.5 (9.9 - 11.2) (n=3) |
| c | 5.3± 0.2 (5.1 - 5.4) (n=3) | 7.3±1.1 (6.3 - 9.1) | 4.8±0.3 (4.5 - 5.2) (n=3) | 4.6 ± 0.5 (4.0 - 5.4) (n=3) |
| c' | 22.1± 1.9 (20 - 24.7) (n=3) | 14.8 ±2.4 (11.2 - 17.1) | 22.3±1.5 (20.3 - 23.7) (n=3) | 24.3±3.2 (20.5 - 28.7) (n=3) |
| V | 34.6± 1.1 (33.4 - 36.1) (n=3) | 33.7 ±2.3 (29.7-35.3) | 34.9±0.8 (33.8 - 35.8) (n=3) | 33.8 ± 0.3 (33.4 - 34.2) (n=3) |
| G1 | 10.3, 9.7 | 10.4±0.6 (9.6 - 11.2) | 11.2± 0.2 (11 - 11.4) (n=3) | 9.1± 0.3 (8.9 - 9.5) (n=3) |
| G2 | 10.2 ± 0.4 (9.8 - 10.7) | 11.1 ±1.2 (9.8 - 13.0) | 11.4± 0.6 (10.5 - 12) (n=3) | 9.3 ± 0.1 (9.2 - 9.5) (n=3) |
| Lip region width | 12.3±0.5 (12 - 13) | 11.2 ±0.4 (11 - 12) | 11 | 10.4± 0.5 (10 - 11) |
| Lip region height | 5 | 4.5±0.5 (4 - 5) | 4.6±0.5 (4 - 5) | 4.7 ± 0.4 (4 - 5) |
| Amphid aperture width | 6.6±0.4 (6 - 7) | 5 | 5.2 ±0.4 (5 - 6) | 6 |
| Odontostyle length | 7 | 7.3±0.4 (7 - 8) | 7 | 6.8 ±0.2 (6 - 7) |
| Odontophore length | 17±0.5 (16 - 18) | 17.5±0.5 (17 - 18) | 16.5±1.1 (15 - 18) | 15± 1.9 (12 - 17) |
| Guiding ring from ant. End | 8.5±0.8 (8 - 10) | 8 | 8±0.9 (7 - 9) | 7.2± 1.0 (6 - 9) |
| Nerve ring from ant. End | 146.1±3.3 (142 - 151) | 137.7 ±2.3 (135 - 140) | 141±1.4 (140 - 144) | 126± 5.7 (115 - 130) |
| Neck length | 328±16.6 (303 - 348) | 318.5±2.9 (314 - 321) | 331.6± 14.1 (319 - 358) | 304.8± 6.2 (294 - 311) |
| Expanded part of pharynx | 104.3±9.1 (86 - 114) | 110±6.1 (105 - 120) | 113±12.1(101 - 135) | 114.8± 8.8 (105 - 130) |
| Cardia length | 15.3±1.5 (14 - 18) | 16.7 ±0.8 (16 - 18) | 13.6±1.9 (11 - 16) | 15.8± 1.0 (15 - 17) |
| Anterior genital branch | 367.2±27.2 (341 - 418) | 354.2±24.3 (330 - 381) | 375±21.4 (341 - 396) | 298± 12.1 (274 - 306) |
| Posterior genital branch | 374.8±37.6 (314 - 434) | 378.2±42.7 (336 - 442) | 375±20.4 (345 - 406) | 308.4± 13 (284 - 318) |
| Vaginal depth | 36.3±1.5 (34 - 38) | 34.5±0.5 (34 - 35) | 34.4±0.5 (34 - 35) | 31.7 ± 1.3 (30 - 33) |
| Vulva from ant. end | 1296.8± 84 (1161 - 1413) | 1144.4±76 (1019 - 1226) | 1197.4±115.6 (1063 - 1382) | 1095± 62.9 (1024 - 1167) |
| Prerectum length | 442.6±24.1 (408 - 475) | 420.7±34.4 (396 - 480) | 385.6±61.4 (283 - 448) | 384± 56.1 (298 - 435) |
| Rectum length | 40.6±1.6 (38 - 43) | 43.2 ±1.9 (41 - 46) | 43±2.4 (40 - 45) | 40.2± 3.2 (35 - 45) |
| Tail length | 736± 40.9 (695 - 792) (n=3) | 469.7±66.2 (370 - 550) | 694±62.5 (610 - 760) (n=3) | 750.6± 44.1 (697 - 805) (n=3) |
| broken L= (230 - 623) | (broken L) | broken L = (495,560) | broken L =620,635 |
Measurements of Paraoxydirus indicus n_ sp_ (All the measurements in μm except L in mm)_
| Locality | Type population | Kerala population | ||
|---|---|---|---|---|
| Characters | Holotype female | Paratype females | Paratype males | Female |
| n | 3 | 9 | 1 | |
| L | 4.0 | 4.0 ± 0.4 (3.6 – 4.6) | 3.8 ± 0.1 (3.7 – 4.0) (n = 4) | 4.8 |
| Body width at neck base | 48 | 50 ± 2.2 (48 – 53) | 47 ± 2.4 (44 – 51) | 60 |
| Body width at mid body | 56 | 58 ± 2.4 (55 – 61) | 49.1 ± 3.7 (44 – 55) | 67 |
| Body width at anus/ cloaca | 30 | 31 ± 1.4 (30 – 33) | 33.8 ± 1.8 (31 – 36) | 32 |
| a | 72.8 | 70.4 ± 4.5 (66.1 – 76.6) | 78.7 ± 4.2 (72.8 – 82.2) (n = 4) | 71.5 |
| b | 12.2 | 11.2 ± 0.7 (10.4 – 12.2) | 11.5 ± 0.6 (11 – 12.5) (n = 4) | 13.2 |
| c | 5.4 | 6.3 ± 0.4 (5.85 – 6.6) | 5.4 ± 0.2 (5.2 – 5.7) (n = 4) | 5.4 |
| c’ | 24.9 | 20.7 ± 0.5 (20 – 21.3) | 21.2 ± 1.5 (19.4 – 23.3) (n = 4) | 27.9 |
| V | 32.1 | 32.8 ± 1.1 (31.2 – 33.6) | - | 30.6 |
| G1 | 10.5 | 10.1 ± 0.4 (9.7 – 10.7) | - | 9.3 |
| G2 | 12.7 | 12.1 ± 1.4 (11 – 14) | - | 11.3 |
| Lip region width | 13 | 13.6 ± 0.5 (13 – 14) | 12.8 ± 0.6 (12 – 14) | 12 |
| Lip region height | 5 | 5 ± 0.8 (4 – 6) | 5.4 ± 0.5 (5 – 6) | 5 |
| Amphid aperture | 8 | 8.3 ± 0.5 (8 – 9) | 7.6 ± 0.8 (7 – 9) | |
| Odontostyle length | 7 | 7.5 ± 0.4 (7 – 8) | 7.4 ± 0.4 (7 – 8) | 7 |
| Odontophore length | 17 | 17.6 ± 0.5 (17 – 18) | 17.3 ± 1.0 (16 – 19) | 17 |
| Guiding ring from ant. End | 9 | 9.3 ± 0.5 (9 – 10) | 8.3 ± 0.5 (8 – 9) | 8 |
| Nerve ring from ant. End | 153 | 149.3 ± 6.0 (141 – 155) | 146.5 ± 2.1 (142 – 149) | 153 |
| Neck length | 334 | 361.6 ± 14.3 (347 – 381) | 344.7 ± 15.4 (314 – 366) | 362 |
| Expanded part of pharynx | 107 | 122.6 ± 6.6 (114 – 130) | 114.3 ± 10.3 (102 – 129) | 146 |
| Cardia length | 25 | 24 ± 2.2 (21 – 26) | 22 ± 2.4 (17 – 25) | 25 |
| Anterior genital branch | 429 | 415 ± 27.7 (388 – 453) | - | 445 |
| Posterior genital branch | 519 | 503 ± 114.2 (404 – 663) | - | 542 |
| Vaginal depth | 34 | 31.6 ± 1.2 (30 – 33) | - | 35 |
| Vulva from ant. End | 1310 | 1339 ± 100 (1217 – 1463) | - | 1464 |
| Prerectum length | 519 | 545.3 ± 41.9 (500 – 601) | 532.5 ± 79.8 (412 – 660) | 597 |
| Rectum length | 41 | 41.6 ± 0.5 (41 – 42) | 46.7 ± 2.8 (44 – 52) | 41 |
| Tail length | 748 | 642 ± 44.3 (602 – 704) (broken L) | 704 ± 26.9 (671 – 746) (n = 4) broken L (565 – 680) | 894 |
| Spicule length | - | - | 49.5 ± 2.2 (45 – 53) | - |
| Lateral guiding pieces | - | - | 13.7 ± 1.5 (12 – 16) | - |
| Ventromedian supplements | - | - | 12 – 15 | - |
Compendium of morphometrics of species belonging to the genus Paraoxydirus Jairajpuri & Ahmad, 1978 (Measurements in μm except L in mm)
| Character species | n | L | a | b | c | c' | v | Lrd | Odont. | Neck | Ph.exp. | Abd | Prerect. | Tail length | Vms | Spicule length | Geo. Dist. | References |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| abnormus | 4♀♀ | 5.0-6.0 | 102-124 | 13-18 | 8-11 | 18-20 | 35-39 | 12 | 8 | 342-416 | 118-146 | 31-34 | 279-331 | 596-668 | - | - | Karnataka- India | Dhanam & Jairajpuri, 1998 |
| 3♂♂ | 5.0-6.0 | 111-130 | 14-16 | 12-16 | - | - | - | 8 | 350-365 | 113-129 | 36-37 | 566-681 | 354-465 | 12-15 | 48-53 | |||
| cavenessi | 5♀♀ | 1.9-4.0 | 59-78 | 6.5-11.7 | 5-10 | 13-21 | 35-44 | 10 | 5-7 | 252-350 | 95-140 | 19.4-28 | 175-311 | 252-588 | - | - | Nigeria | Ferris et al., 1980 |
| 3♂♂ | 2.0-2.5 | 63-66 | 6.4-9.0 | 5-8 | - | - | - | - | 280-315 | 100-144 | - | - | 336-448 | 7-8 | 36-37 | |||
| gigas | 2♀♀ | 3.8-4.5 | 65-66 | 12-13 | 10 | - | 40-42 | 11* | 5* | - | - | - | - | - | - | - | Nainital- India | Jairajpuri, 1964 |
| 233 | 3.7-4.3 | 61-86 | 12-13 | 11-12 | 13 | - | 11* | 5* | - | - | 30* | 300* | 390* | 15 | 50 | |||
| 1♂ | 3.7 | 88 | 13 | 14 | - | - | - | 6 | - | - | - | - | 15 | 49 | M.P.- India | Siddiqi, 1966 | ||
| 1♀ | 3.9 | 70 | 13 | 9 | - | 39 | - | 5-7 | - | - | - | - | - | - | - | UP-India | Baqri & Jairajpuri, 1968 | |
| 233 | 3.6-3.7 | 84-86 | 12.0-12.2 | 10-15 | - | - | - | - | - | - | - | - | - | 13 | 48-51 | |||
| 25♀♀ | 3.7-4.3 | 72-83 | 11.2-12.9 | 11-17 | 14* | 42-45 | 12* | 6-7 | 320-350 | 140* | 28-34 | 260-360 | 250-345 | - | - | Several localities-India | Jairajpuri & Ahmad, 1978 | |
| 10♀♀ | 3.7-4.0 | 66-79 | 11.5-12.9 | 10-16 | 13* | - | - | 6-7 | 290-305 | 33-35 | - | 220-350 | 11-13 | 50-55 | ||||
| 8♀♀ | 4.1-4.6 | 77-85 | 13-15 | 7-8 | 18-21 | 35-42 | 10-11 | 5-6 | 304-326 | 93-108 | 28-31 | 260-303 | 425-604 | - | - | Goa- India | Present paper | |
| 5♀♀ | 3.8-4.5 | 80-88 | 11-14 | 8-10 | 12-14 | - | 10-11 | 5 | 309-342 | 88-104 | 33-36 | 370-457 | 440-480 | 10-14 | 52-54 | |||
| 4♀ | 3.3-4.5 | 64.4-73 | 11-13.7 | 9-11 | 10.7-15.4 | 36.3-40.4 | 10 | 5-6 | 302-341 | 85-118 | 28-32 | 220-319 | 300-478+ | - | - | Assam-India | Present paper | |
| 1♂ | 3.5 | 71.3 | 10.7 | 10.6 | 10.3 | - | 10 | 6 | 325 | 113 | 32 | 280 | 330+ | 12 | 57 | |||
| magnus | 6♀♀ | 4.8-5.7 | 85-117 | 13-17 | 12-15 | 12.5 | 33-40 | 10 | 8 | 334-367 | 84-118 | 32.8 | 230 | 389-400 | - | - | Bangladesh | Timm, 1964 |
| 6♂♂ | 4.4-4.9 | 87-103 | 14-15.4 | 17- 20 | - | - | - | - | - | - | - | - | - | 12-16 | 54-57 | |||
| novus | 2♀♀ | 3.9-4.0 | 65-68 | 11-12 | 4-6 | 20 | 34-35 | 10** | 7 | - | - | 33* | 17abd | 615* | - | - | Andaman- India | Jairajpuri, 1965 |
| 5♀♀ | 3.1-3.4 | 53-68 | 10-11 | 4.0-5.4 | 21-29 | 33-34 | 10-11 | 6-7 | 294-311 | 105-130 | 27-31 | 298-435 | 697-805 | - | - | Assam-India | Present paper | |
| 5♀♀ | 3.2-3.5 | 52-58 | 10-11 | 4.5-5.0 | 20-24 | 34-36 | 11 | 7 | 319-358 | 101-135 | 30-33 | 283-448 | 610-760 | - | - | Goa-India | Present paper | |
| 6♀♀ | 3.7-4.0 | 57-66 | 11 -12 | 5.1-5.4 | 20-25 | 33-36 | 12-13 | 7 | 303-348 | 86-114 | 31-36 | 408-475 | 695-792 | - | - | Tamil Nadu | Present paper | |
| 4♀♀ | 3.3-3.5 | 55-66 | 10-11 | 6.3-9.0 | 11-17 | 30-35 | 11-12 | 7-8 | 314-321 | 105-120 | 30-33 | 396-480 | 370-550+ | - | - | Karnataka-India | Present paper | |
| pelinus | 3♀♀ | 5.2-5.4 | 102-140 | 17-20 | 9-11 | 20 | 29-34 | 9 | 7 | 266-311 | 90-96 | 26 | 112-256 | 496-592 | - | - | Philipines | Ferris et al., 1980 |
| 2♂♂ | 4.8-4.85 | 89-127 | 15.0-15.4 | 12-17 | - | - | - | - | 311 | 96 | - | - | 281-383 | 11-13 | 50-55 | |||
| indicus sp.n. | 4♀♀ | 3.6-4.6 | 66-76 | 10.5-12.3 | 5.4-6.6 | 21-25 | 31-33 | 13-14 | 7-8 | 334-381 | 107-130 | 30-33 | 500-601 | 748 (602-704+) | - | - | Tamil Nadu-India | Present paper |
| 9♂♂ | 3.7-4.0 | 73-82 | 11.0-12.5 | 5.2-5.7 | 19-23 | - | 12-14 | 7-8 | 314-366 | 102-129 | 31-36 | 412-600 | 671-746 | 12-15 | 45-53 | |||
| 1♀ | 4.8 | 71.5 | 13.2 | 5.4 | 28 | 30.6 | 12 | 7 | 362 | 146 | 32 | 597 | 894 | - | - | Karnataka-India | Present paper | |
| vulvalpapillatus sp.n. | 6♀♀ | 4.3-4.8 | 83-95 | 13-15 | 13-14 | 10-13 | 36-39 | 10-11 | 5-6 | 317-352 | 82-107 | 25-32 | 315-435 | 325 (310-365+) | - | - | Goa-India | Present paper |
| 8♂♂ | 3.8-5.0 | 69-93 | 13.0-15.2 | 8-20 | 7-15 | - | 10-11 | 4-5 | 297-339 | 76-107 | 32-39 | 432-602 | 247-475 | 11-14 | 46-54 | |||
| 1♀ | 4.65 | 72 | 13 | 15 | 9.8 | 42 | 10 | 5 | 352 | 138 | 31 | 290 | 312 | - | - | Karnataka-India | Present paper |
Measurements of Paraoxydirus gigas Jairajpuri, 1964 (All the measurements in μm except L in mm)_
| Locality | Goa population | Assam population | Nainital (type population) | Bijnore population (After Baqri & Jairajpuri, 1968) | others | |||||
|---|---|---|---|---|---|---|---|---|---|---|
| Characters | Females | Males | Females | Male | Females | Males | Female | Males | Females | Males |
| n | 8 | 5 | 4 | 1 | 2 | 2 | 1 | 2 | 25 | 10 |
| L | 4.3 ± 0.2 (4.1 - 4.6) | 4.2 ± 0.3 (3.8 - 4.5) | 3.9 ± 0.4 (3.3 - 4.5) | 3.5 | 3.8 - 4.5 | 3.7 - 4.3 | 3.89 | 3.6, 3.7 | 3.7 - 4.3 | 3.7 - 4.0 |
| Body width at neck base | 44.2 ± 2.2 (42 - 49) | 42.2 ± 1.0 (42 - 45) | 46.2 ± 3.3 (42 - 51) | 40 | 50* | - | - | 50* | - | |
| Body width at mid body | 54.4 ± 2.7 (50 - 59) | 50.6 ± 2.3 (48 - 54) | 57.2 ± 4.8 (52 - 62) | 49 | - | - | - | - | ||
| Body width at anus/ cloaca | 30 ± 1.1 (28 - 31) | 34.7 ± 1.1 (33 - 36) | 30.7 ± 1.6 (28 - 32) | 32 | 30* | - | - | 28 - 34 | 33 - 35 | |
| a | 81.4 ± 3.2 (77 - 85.3) | 83 ± 2.7 (80.5 - 87.7) | 69.2 ± 3.4 (64.4 - 73) | 71.3 | 65 - 66 | 61 - 86 | 70 | 84, 86 | 72 - 83 | 66 - 79 |
| b | 14 ± 0.5 (13.5 - 14.7) | 13.1 ± 1.0 (11.3 - 14.1) | 12.3 ± 0.9 (11.1 - 13.7) | 10.7 | 12 - 13 | 12 - 13 | 12.8 | 12, 12.2 | 11 - 13 | 11.5 - 13 |
| c | 7.6 ± 0.3 (7.2 - 8.2) | 9.2 ± 0.8 (8.2 - 10.3) | 9.8 ± 0.8 (9.1 - 11.2) | 10.6 | 10 | 11 - 12 | 9 | 10, 15 | 11 - 17 | 10 - 16 |
| c' | 19.2 ± 0.8 (18.2 - 20.6) | 13.2 ± 0.7 (12.2 - 13.9) | 13.2 ± 1.7 (10.7 - 15.4) | 10.3 | 13 | 12 | - | - | 14* | 13.5* |
| V | 38.7 ± 2.4 (34.8 - 42.1) | - | 37.8 ± 1.5 (36.3 - 40.4) | - | 40 - 42 | - | 39 | - | 42 - 45 | - |
| G1 | 8.2 ± 1.0 (6.7 - 9.3) | - | 11.3 ± 1.2 (9.8 - 13) | - | - | - | - | - | 9 - 15 | - |
| G2 | 8.3 ± 0.8 (7.4 - 9.6) | - | 12.5 ± 1.3 (10.6 - 14) | - | - | - | - | - | 9 - 18 | - |
| Lip region width | 10.2 ± 0.4 (10 - 11) | 10.4 ± 0.5 (10 - 11) | 10 | 10 | - | 11* | - | - | 11* | - |
| Lip region height | 4.5 ± 0.5 (4 - 5) | 4.2 ± 0.4 (4 - 5) | 4 | 5 | - | 5* | - | - | 4* | |
| Amphid aperture width | 6.5 ± 0.5 (6 - 7) | 6.5 ± 0.5 (6 - 7) | 6.2 ± 0.4 (6 - 7) | 7 | - | 6* | - | - | - | - |
| Odontostyle length | 5.1 ± 0.3 (5 - 6) | 5 | 5.3 ± 0.5 (5 - 6) | 6 | - | 5* | - | 5, 7 | 6 - 7 | 6 - 7 |
| Odontophore length | 12.2 ± 0.6 (11 - 13) | 13 | 10.6 ± 0.9 (10 - 12) | 11 | - | 10* | - | 8, 10 | 10 - 12 | 12 - 13 |
| Guiding ring from ant. End | 5 | 5 | 5.3 ± 0.5 (5 - 6) | 6 | - | - | - | - | 6* | - |
| Nerve ring from ant. End | 145.5 ± 7.1 (137 - 157) | 148.2 ± 5.3 (141 - 157) | 144.2 ± 1.9 (142 - 147) | 140 | - | - | - | - | 130* | |
| Neck length | 314.7 ± 7.2 (304 - 326) | 321.8 ± 11.2 (309 - 342) | 321.2 ± 15.5 (302 - 341) | 325 | - | - | - | - | 320 - 350 | 290 - 305 |
| Expanded part of pharynx | 101 ± 4.2 (93 - 108) | 94.4 ± 5.8 (88 - 104) | 101.5 ± 13.5 (85 - 118) | 113 | - | - | - | - | 130* | - |
| Cardia length | 14.3 ± 1.6 (12 - 17) | 16.2 ± 0.8 (15 - 17) | 17.2 ± 3.3 (13 - 21) | 17 | - | 14* | - | - | 20* | - |
| Anterior genital branch | 364.3 ± 38 (305 - 405) | - | 447.2 ± 60.6 (398 - 547) | - | - | - | - | - | - | - |
| Posterior genital branch | 372.4 ± 31.6 (320 - 410) | - | 495.2 ± 66 (405 - 590) | - | - | - | - | - | - | - |
| Vaginal depth | 29.4 ± 2.3 (27 - 34) | - | 31, 32 | - | - | - | - | - | 38* | - |
| Vulva from ant. end | 1708 ± 94 (1519 - 1795) | - | 1504.3 ± 199 (1242 - 1698) | - | - | - | - | - | - | - |
| Prerectum length | 277.1 ± 34.4 (260 - 303) | 418 ± 29.3 (370 - 457) | 271 ± 35.1 (220 - 319) | 280 | - | - | - | - | 260 - 360 | - |
| Rectum length | 34.5 ± 3.1 (30 - 39) | 51.5 ± 2.7 (48 - 55) | 37, 41 | 50 | - | 50* | - | - | 29 - 36 | - |
| Tail length | 577.5 ± 13.3 (565 - 604) | 454.6 ± 14.6 (440 - 480) | 407.5 ± 65.9 (300 - 478) | - | - | - | - | 250 - 345 | 220 - 350 | |
| (broken) | broken | |||||||||
| Spicule length | - | 52.7 ± 0.8 (52 - 54) | - | 57 | - | 50 | - | 48, 51 | - | 50 - 55 |
| Lateral guiding pieces | - | 11.2 ± 1.5 (9 - 13) | - | 12 | - | - | - | - | - | 10 - 11 |
| Ventromedian supplements | - | 10 - 14 | - | 11 | - | 15 | - | 13 | - | 11 - 13 |