References
- Aldrich, P. R., Parker, G. R., Romero-Severson, J., Michler, C. H., 2005: Confirmation of oak recruitment failure in Indiana old-growth forest: 75 years of data. Forest Science, 51:406–416.
- Bauhus, J., Puettmann, K., Messier, C., 2009: Silviculture for old-growth attributes. Forest Ecology and Management, 258:525–537.
- Begović, K., Schurman, J. S., Svitok, M., Pavlin, J., Lang-behn, T., Svobodová, K. et al., 2023: Large old trees increase growth under shifting climatic constraints: Aligning tree longevity and individual growth dynamics in primary mountain spruce forests. Global Change Biology, 29:143–164.
- Cerioni, M., Brabec, M., Bače, R., Bāders, E., Bončina, A., Brůna, J. et al., 2024: Recovery and resilience of European temperate forests after large and severe disturbances. Global Change Biology, 30:e17159.
- Čada, V., Svoboda, M., Janda, P., 2013: Dendrochrono-logical reconstruction of the disturbance history and past development of the mountain Norway spruce in the Bohemian Forest, Central Europe. Forest Ecology and Management, 295:59–68.
- Čada, V., Morrisey, R. C., Michalová, Z., Bače, R., Janda, P., Svoboda, M., 2016: Frequent severe natural disturbances and non-equilibrium landscape dynamics shaped the mountain spruce forest in central Europe. Forest Ecology and Management, 363:169–178.
- Červenka, J., Bače, R., Zenáhlíková, J., Svoboda, M., 2016: Změny porostní struktury, množství a kvality mrtvého dřeva v horské smrčině po velkoplošné disturbanci. Zprávy lesnického výzkumu, 61:254–261. (In Czech).
- Diaci, J., Rozenbergar, D., Boncina, A., 2010: Stand dynamics of Dinaric old-growth forest in Slovenia: Are indirect human influences relevant? Plant Bio-systems, 144:194–201.
- Diaci, J., Rozenbergar, D., Anic, I., Mikac, S., Saniga, M., Kucbel, S. et al., 2011: Structural dynamics and synchronous silver fir decline in mixed old-growth mountain forests in Eastern and Southeastern Europe. Forestry, 84:479–491.
- Di Filippo, A., Biondi, F., Piovesan, G., Ziaco, E., 2017: Tree ring-based metrics for assessing old-growth forest naturalness. Journal of Applied Ecology, 54:737–749.
- Goff, F. G., West, D., 1975: Canopy-understory interaction effects on forest population structure. Forest Science, 21:98–108.
- Goodburn, J. M., Lorimer, C. G., 1999: Population structure in old-growth and managed northern hard-woods: an examination of the balanced diameter distribution concept. Forest Ecology and Management, 118:11–29.
- Havira, M., Bače, R., Čada, V., Svoboda, M., 2017: Množství a výšková struktura odrostlého zmlazení v horském smrkovém lese Hrubého Jeseníku. Zprávy lesnického výzkumu, 62:33–41. (In Czech).
- Heiri, C., Wolf, A., Rohrer, L., Bugmann, H., 2009: Forty years of natural dynamics in Swiss beech forests: structure, composition, and the influence of former management. Ecological Applications, 19:1920–1934.
- Holeksa, J., 2001: Coarse woody debris in a Carpathian subalpine spruce forest. Forstwissenschaftliches Centralblatt, 120:256–270.
- Holeksa, J., Saniga, M., Szwagrzyk, J., Dziedzic, Ferenc, S., Wodka, M., 2007: Altitudinal variability of stand structure and regeneration in the subalpine spruce forests of the Poľana biosphere reserve, Central Slovakia. European Journal of Forest Research, 126:303–313.
- Holeksa, J., Zielonka, T., Źywiec, M., Fleischer, P., 2016: Identifying the disturbance history over a large area of larch–spruce mountain forest in Central Europe. Forest Ecology and Management, 361:318–327.
- Holeksa, J., Jaloviar, P., Kucbel, S., Saniga, M., Svoboda, M., Szewczyk, J. et al., 2017: Models of disturbance driven dynamics in the West Carpathian spruce forests. Forest Ecology and Management, 388:79–89.
- Holeksa, J., Źywiec, M., Bogdziewicz, M., Kurek, P., Milne-Rostkowska, F., Piechnik, Ł. et al., 2021: Microsite-specific 25-year mortality of Norway spruce saplings. Forest Ecology and Management, 498:119572.
- Janda, P., Svoboda, M., Bače, R., Čada, V., Peck, J. L. E., 2014: Three hundred years of spatio-temporal development in a primary mountain Norway spruce stand in the Bohemian Forest, central Europe. Forest Ecology and Management, 330:304–311.
- Janda, P., Trotsiuk, V., Mikoláš, M., Bače, R., Nagel, T. A., Seidl, R. et al., 2017: The historical disturbance regime of mountain Norway spruce forests in the Western Carpathians and its influence on current forest structure and composition. Forest Ecology and Management, 388:67–78.
- Janda, P., Tepley, A. J., Schurman, J. S., Brabec, M., Nagel, T. A., Bače, R. et al., 2019: Drivers of basal area variation across primary late-successional Picea abies forests of the Carpathian Mountains. Forest Ecology and Management, 435:196–204.
- Korpeľ, Š., 1989: Pralesy Slovenska. Bratislava, Veda, 332 p. (In Slovak).
- Kräuchi, N., Brang, P., Schönenberger, W., 2000: Forests of mountainous regions: gaps in knowledge and research needs. Forest Ecology and Management, 132:73–82.
- Kucbel, S., Saniga, M., Jaloviar, P., Vencurik, J., 2012: Stand structure and temporal variability in old-growth beech-dominated forests of the northwestern Carpathians: A 40-years perspective. Forest Ecology and Management, 264:125–133.
- Kucbel, S., 2014: Časová a priestorová variabilita štruk-Časová a priestorová variabilita štruktúry vysokohorských smrekových lesov Slovenska. Zvolen, Technická univerzita vo Zvolene, 101 p. (In Slovak).
- Lamedica, S., Lingua, E., Popa, I., Motta, R., Carrer, M., 2011: Spatial structure in four Norway spruce stands with different management history in the Alps and Carpathians. Silva Fennica, 45:865–873.
- Lausch, A., Heurich, M., Fahse, L., 2013: Spatio-temporal infestation patterns of Ips typographus (L.) in the Bavarian Forest National Park, Germany. Ecological Indicators, 31:73–81.
- Leak, W. A., 1996: Long-term structural change in uneven-aged northern hardwoods. Forest Science, 42:160–165.
- Leibundgut, H., 1982: Europäische Urwälder der Berg-Europäische Urwälder der Bergstufe. Bern, Verlag Paul Haupt, 308 p. (In German).
- Leibundgut, H., 1993: Europäische Urwälder. Bern, Verlag Paul Haupt, 260 p. (In German). Lindenmayer, D. B., 2017: Conserving large old trees as small natural features. Biological Conservation, 211:51–59.
- Lindenmayer, D. B., Laurance, W. F., 2017: The ecology, distribution, conservation and management of large old trees. Biological Reviews, 92:1434–1458.
- Lorimer, C. G., 1980: Age structure and disturbance history of a southern Appalachian virgin forest. Ecology, 61:1169–1184.
- Lukasová, V., Škvareninová, J., Bičárová, S., Sitárová, Z., Hlavatá, H., Borsányi, P. et al., 2021: Regional and altitudinal aspects in summer heatwave intensification in the Western Carpathians. Theoretical and Applied Climatology, 146:1111–1125.
- Marini, L., Matthew, P. A., Battisti, A., Faccoli, M., 2012: Climate affects severity and altitudinal distribution of outbreaks in an eruptive bark beetle. Climatic Change, 115:327–341.
- Mayer, H., Ott, E., 1991: Gebirgswaldbau – Schutzwald-Gebirgswaldbau – Schutzwaldpflege. Stuttgart, Gustav Fischer Verlag, 587 p. (In German).
- Mezei, P., Grodzki, W., Blaženec, M., Jakuš, R., 2014: Factors influencing the wind–bark beetles’ disturbance system in the course of an Ips typographus outbreak in the Tatra Mountains. Forest Ecology and Management, 312:67–77.
- Mikoláš, M., Ujházy, K., Jasík, M., Wiezik, M., Gallay, I., Polák, P. et al., 2019: Primary forest distribution and representation in a Central European landscape: Results of a large-scale field-based census. Forest Ecology and Management, 449:11–18.
- Nováková, M., Edwards-Jonášová, M., 2015: Restoration of Central-European mountain Norway spruce forest 15 years after natural and anthropogenic disturbance. Forest Ecology and Management, 344:120–130.
- Orman, O., Adamus, M., Foremnik, K., 2023: Norway spruce sapling plasticity in their responses of architecture and growth to light gradient decreases with altitude in subalpine stands. Forest Ecology and Management, 536:120898.
- Ott, E., Frehner, M., Frey, H., Lüscher, P., 1997: Gebirgsnadelwälder. Bern, Verlag Paul Haupt, 287 p. (In German).
- Petráš, R., Pajtík, J., 1991: Sústava československých objemových tabuliek drevín. Lesnícky časopis, 37:49–56. (In Slovak).
- Primicia, I., Camarero, J. J., Janda, P., Čada, V., Morrissey, R. C., Trotsiuk, V. et al. 2015: Age, competition, disturbance and elevation effects on tree and stand growth response of primary Picea abies forest to climate. Forest Ecology and Management, 354:77–86.
- Prodan, M., 1951: Messung der Waldbestände. Frank-Frank-furt am Main, Sauerländer’s Verlag, 260 p. (In Ger-260 p. (In German).
- Schmidt-Vogt, H., 1991: Die Fichte, Bd. II/3. Hamburg, Berlin, Paul Parey Verlag, 781 p. (In German).
- Schönenberger, W., 1986: Rottenaufforstung im Gebirge. Schweizerische Zeitschrift für Forstwesen, 137:501–509. (In German).
- Schurman, J. S., Trotsiuk, V., Bače, R., Čada, V., Fraver, S., Janda, P., 2018: Large-scale disturbance legacies and the climate sensitivity of primary Picea abies forests. Global Change Biology, 24:2169–2181.
- Schurman, J. S., Janda, P., Rydval, M., Mikolaš, M., Svoboda, M., Babst, F., 2024: Climate-competition tradeoffs shape the range limits of European beech and Norway spruce along elevational gradients across the Carpathian Mountains. Ecography:e06715.
- Seidl, R., Thom, D., Kautz, M., Martin-Benito, D., Peltoniemi, M., Vacchiano, G., 2017: Forest disturbances under climate change. Nature Climate Change, 7:395–402.
- Spinu, A. P., Petritan, I. C., Mikoláš, M., Janda, P., Vostarek, O., Čada, V. et al., 2020: Moderate- to high-severity disturbances shaped the structure of primary Picea abies [L.] Karst. forest in the Southern Carpathians. Forests, 11:1315.
- Svoboda, M., Pouska, V., 2008: Structure of a Central-European mountain spruce old-growth forest with respect to historical development. Forest Ecology and Management, 255:2177–2188.
- Svoboda, M., Fraver, S., Janda, P., Bače, R., Zenáhlíková, J., 2010: Natural development and regeneration of a Central European montane spruce forest. Forest Ecology and Management, 260:707–714.
- Svoboda, M., Janda, P., Nagel, T. A., Fraver, S., Rejzek, J., Bace, R., 2012: Disturbance history of an old-growth sub-alpine Picea abies stand in the Bohemian Forest, Czech Republic. Journal of Vegetation Science, 23:86–97.
- Svoboda, M., Janda, P., Bače, R., Fraver, S., Nagel, T. A., Rejzek, J. et al., 2014: Landscape-level variability in historical disturbance in primary Picea abies mountain forests of the Eastern Carpathians, Romania. Journal of Vegetation Science, 25:386–401.
- Svobodová, K., Langbehn, T., Björklund, J., Rydval, M., Trotsiuk, V., Morrissey, R. C. et al., 2019: Increased sensitivity to drought across successional stages in natural Norway spruce (Picea abies [L.] Karst.) forests of the Calimani Mountains, Romania. Trees, 33:1345–1359.
- Synek, M., Janda, P., Mikoláš, M., Nagel, T. A., Schurman, J. S., Pettit, J. L. et al., 2020: Contrasting patterns of natural mortality in primary Picea forests of the Carpathian Mountains. Forest Ecology and Management, 457:117734.
- Škvarenina, J., Tomlain, J., Hrvoľ, J., Škvareninová, J., Nejedlík, P., 2009: Progress in dryness and wetness parameters in altitudinal vegetation stages of West Carpathians: Time-series analysis 1951–2007. Idojárás, 113:47–54.
- Šmelko, Š., 2000: Dendrometria. Zvolen, Technická univerzita vo Zvolene, 399 p. (In Slovak).
- Trotsiuk, V., Svoboda, M., Janda, P., Mikolas, M., Bace, R., Rejzek J. et al., 2014: A mixed severity disturbance regime in the primary Picea abies [L.] Karst. forests of the Ukrainian Carpathians. Forest Ecology and Management, 334:144–153.
- Vencurik, J., Kucbel, S., Saniga, M., Jaloviar, P., Sedmáková, D., Pittner, J. et al., 2020: Growth dynamics of the Norway spruce and silver fir understory in continuous cover forestry. iForest, 13:56–64.
- Vrška, T., Adam, D., Hort, L., Kolář, T., Janík, D., 2009: European beech (Fagus sylvatica L.) and silver fir (Abies alba Mill.) rotation in the Carpathians – A developmental cycle or a linear trend induced by man? Forest Ecology and Management, 258:347–356.
- Wasser, B., Frehner, M., 1996: Wegleitung Minimale Pflegemassnahmen für Wälder mit Schutzfunktion. Bern, Bundesamt für Umwelt,Wald und Landschaft, 122 p. (In German).
- Weigel, R., Muffler, L., Klisz, M., Kreyling, J., van der Maaten-Theunissen, M., Wilmking, M. et al., 2018: Winter matters: sensitivity to winter climate and cold events increases towards the cold distribution margin of European beech (Fagus sylvatica L.). Journal of Biogeography, 45:2779–2790.
- Westphal, C., Tremer, N., von Oheimb, G., Hansen, J., von Gadow, K., Härdtle, W., 2006: Is the reverse J-shaped diameter distribution universally applicable in European virgin beech forests? Forest Ecology and Management, 223:75–83.
- Winter, M. B., Baier, R., Ammer, C., 2015: Regeneration dynamics and resilience of unmanaged mountain forests in the Northern Limestone Alps following bark beetle-induced spruce dieback. European Journal of Forest Research, 134:949–968.
- Woods, K. D., Nagel, T. A., Brzeziecky, B., Cowell, C. M., Firm, D., Jaloviar, P. et al., 2021: Multi-decade tree mortality in temperate old-growth forests of Europe and North America: Non-equilibrial dynamics and species-individualistic response to disturbance. Global Ecology and Biogeography, 30:1311–1333.
- Zhang, L., Gove, J. H., Liu, C., Leak, W. B., 2001: A finite mixture of two Weibull distributions for modeling the diameter distributions of rotated-sigmoid, uneven-aged stands. Canadian Journal of Forest Research, 31:1654–1659.
- Zielonka, T., 2006: Quantity and decay stages of coarse woody debris in old-growth subalpine spruce forests of the western Carpathians, Poland. Canadian Journal of Forest Research, 36:2614–2622.
- Zielonka, T., Holeksa, J., Fleischer, P., Kapusta, P., 2010: A tree-ring reconstruction of wind disturbances in a forest of the Slovakian Tatra Mountains, Western Carpathians. Journal of Vegetation Science, 21:31–42.
- Zlatník, A., 1976: Lesnická fytocenologie. Praha, SZN, 495 p. (In Slovak).
- Źywiec, M., Ledwoń, M., 2008: Spatial and temporal patterns of Rowan (Sorbus aucuparia L.) regeneration in subalpine spruce forest. Plant Ecology, 194:283–291.
- Źywiec, M., Holeksa, J., 2012: Sprouting extends the lifespan of tree species in a seedling bank: 12-year study. Forest Ecology and Management, 284:205–212.
- Źywiec, M., Holeksa, J., Wesołowska, M., Szewczyk, J., Zwijacz-Kozica, T., Kapusta, P., 2013: Sorbus aucuparia regeneration in a coarse-grained spruce forest – A landscape scale. Journal of Vegetation Science, 24:735–743.
- Macdonald, P., with contributions from Du, J., 2010: mixdist: Finite Mixture Distribution Models. R package version 0.5–3. http://CRAN.R-project.org/package=mixdist.
- R Development Core Team, 2010: R: A language and environment for statistical computing. R Foundation for Statistical Computing, Vienna, Austria. Available at http://www.R-project.org.