Figure 1
Location of Gomilshanski lisy National Nature Park and sample plots (1 – plot of clear felling in the managed zone; 2 – plot of selective sanitary felling in the managed zone; 3 – protected zone; 4 – zone of regulated recreation; 5 – zone of stationary recreation)
Figure 2
Characteristics of climate in the territory of Gomilshansky lisy National Nature Park (Zmiyiv meteorological station, 49.664N, 36.273E, 112 m a.s.l., climate class DFB. Years: 1990–2019 (Zepner et al. 2020)
Table 1
Location of sample plots
| Sample plots | Latitude [°N] | Longitude [°E] |
|---|---|---|
| 1. Plot of clear felling in the managed zone | 49.5962 | 36.2427 |
| 2. Plot of selective sanitary felling in the managed zone | 49.6204 | 36.2782 |
| 3. Protected zone | 49.6109 | 36.2895 |
| 4. Zone of regulated recreation | 49.6092 | 36.3007 |
| 5. Zone of stationary recreation | 49.6204 | 36.3176 |
Figure 3
Design of window trap (Skrylnik and Bieliavtsev 2020)
Table 2
Xylophagous beetles (Coleoptera) in different zones of Gomilshanski lisy National Nature Park
| Species | Insect counts per groups of sample plots | |||||
|---|---|---|---|---|---|---|
| plot of clear felling in the managed zone | plot of selective sanitary felling in the managed zone | zone of stationary recreation | zone of regulated recreation | protected zone | Total | |
| 1 | 2 | 3 | 4 | 5 | 6 | 7 |
| Curculionidae: Scolytinae | ||||||
| Anisandrus dispar (Fabricius, 1792) | 199 | 87 | 222 | 413 | 426 | 1347 |
| Anisandrus maiche (Stark, 1936) | 5 | 8 | 4 | 0 | 0 | 17 |
| Dryocoetes villosus (Fabricius, 1792) | 0 | 1 | 0 | 0 | 1 | 2 |
| Hylesinus crenatus (Fabricius, 1787) | 0 | 0 | 2 | 1 | 0 | 3 |
| Hylesinus toranio (Danthoine, 1788) | 33 | 0 | 2 | 2 | 14 | 51 |
| Hylesinus varius (Fabricius, 1775) | 6 | 0 | 14 | 0 | 2 | 22 |
| Lymantor coryli (Perris, 1855) | 0 | 1 | 0 | 0 | 0 | 1 |
| Pteleobius vittatus (Fabricius, 1787) | 0 | 0 | 21 | 1 | 0 | 22 |
| Scolytus intricatus (Ratzeburg, 1837) | 2 | 12 | 6 | 0 | 2 | 22 |
| Scolytus koenigi (Schevyrew, 1890) | 2 | 3 | 4 | 7 | 2 | 18 |
| Scolytus laevis (F. Chapuis, 1869) | 0 | 2 | 0 | 0 | 0 | 2 |
| Scolytus mali (Bechstein, 1805) | 1 | 0 | 3 | 0 | 0 | 4 |
| Scolytus multistriatus (Marsham, 1802) | 8 | 18 | 22 | 3 | 1 | 52 |
| Scolytus pygmaeus (Fabricius, 1787) | 0 | 0 | 1 | 0 | 0 | 1 |
| Scolytus rugulosus (Müller, 1818) | 0 | 0 | 4 | 0 | 0 | 4 |
| Scolytus sp. | 0 | 0 | 0 | 2 | 0 | 2 |
| Trypodendron signatum (Fabricius, 1792) | 0 | 1 | 0 | 2 | 4 | 7 |
| Xyleborinus attenuatus (Eichhoff, 1876) | 36 | 18 | 19 | 43 | 73 | 189 |
| Xyleborinus saxesenii (Ratzeburg, 1837) | 701 | 854 | 2053 | 1469 | 2718 | 7795 |
| Xyleborus dryographus (Ratzeburg, 1837) | 5 | 3 | 1 | 4 | 1 | 14 |
| Xyleborus monographus (Fabricius, 1792) | 26 | 10 | 26 | 42 | 87 | 191 |
| Curculionidae: Cossoninae | ||||||
| Rhyncolus ater (Linnaeus, 1758) | 2 | 0 | 1 | 0 | 0 | 3 |
| Cerambycidae | ||||||
| Cerambyx (Microceram byx) scopolii (Fuessly, 1775) | 5 | 1 | 1 | 0 | 7 | |
| Leioderes kollari (Redtenbacher, 1849) | 0 | 3 | 0 | 0 | 0 | 3 |
| Leiopus linnei (Wallin, Nylander, Kvamme, 2009) | 2 | 0 | 0 | 0 | 0 | 2 |
| Mesosa curculionoides (Linnaeus, 1761) | 1 | 0 | 2 | 0 | 0 | 3 |
| Plagionotus arcuatus (Linnaeus, 1758) | 1 | 0 | 0 | 0 | 0 | 1 |
| Plagionotus detritus (Linnaeus, 1758) | 1 | 0 | 0 | 0 | 0 | 1 |
| Pogonocherus hispidulus (Piller, Mitterpacher, 1783) | 2 | 0 | 1 | 2 | 0 | 5 |
| Pyrrhidium sanguineum (Linnaeus, 1758) | 3 | 0 | 3 | 1 | 1 | 8 |
| Rhagium (Megarhagium) mordax (De Geer, 1775) | 1 | 0 | 0 | 0 | 0 | 1 |
| Rhagium sycophanta (Schrank von Paula, 1781) | 0 | 1 | 3 | 1 | 5 | |
| Ropalopus macropus (Germar, 1824) | 0 | 0 | 0 | 1 | 0 | 1 |
| Stenocorus (Anisorus) quercus (Gotz, 1783) | 0 | 1 | 0 | 0 | 0 | 1 |
| Xylotrechus arvicola (Olivier, 1795) | 0 | 1 | 0 | 0 | 0 | 1 |
| Histeridae | ||||||
| Abraeus granulum (Erichson, 1839) | 1 | 1 | 1 | 0 | 4 | 7 |
| Lymexilidae | ||||||
| Elateroides dermestoides (Fleming, 1921) | 2 | 0 | 0 | 1 | 77 | 80 |
| Bostrichidae | ||||||
| Bostrichus capucinus (Linnaeus, 1758) | 0 | 1 | 0 | 1 | 0 | 2 |
| Buprestidae | ||||||
| Agrilus angustulus (Illiger, 1803) | 1 | 0 | 0 | 0 | 0 | 1 |
| Agrilus hastulifer (Ratzeburg, 1839) | 0 | 1 | 0 | 0 | 0 | 1 |
| Agrilus sulcicollis (Boisduval, Lacordaire, 1835) | 1 | 2 | 0 | 0 | 0 | 3 |
| Chrysobothris affinis (Fabricius, 1794) | 0 | 0 | 0 | 1 | 0 | 1 |
| Total individuals | 1047 | 1030 | 2415 | 1997 | 3414 | 9903 |
| Total species | 25 | 22 | 22 | 19 | 16 | 42 |
Figure 4
Number of xylophagous species in different zones of Gomilshanski lisy National Nature Park
Figure 5
Menhinick index of xylophagous insects in different zones of Gomilshanski lisy National Nature Park
Figure 6
Xylophagous species abundance in in different zones of Gomilshanski lisy National Nature Park
Table 3
The proportion of the most abundant xylophagous beetles (Coleoptera) in different zones of Gomilshanski lisy National Nature Park*
| Species | Insect counts per groups of sample plots | |||||
|---|---|---|---|---|---|---|
| plot of clear felling in the managed zone | plot of selective sanitary felling in the managed zone | zone of stationary recreation | zone of regulated recreation | protected zone | Total | |
| Xyleborinus saxesenii | 67.0 | 82.9 | 85.0 | 73.6 | 79.6 | 78.7 |
| Anisandrus dispar | 19.0 | 8.5 | 9.2 | 20.7 | 12.5 | 13.6 |
| Xyleborus monographus | 2.5 | 1.0 | 1.1 | 2.1 | 2.6 | 1.9 |
| Xyleborinus attenuatus | 3.4 | 1.8 | 0.8 | 2.1 | 2.1 | 1.9 |
| Elateroides dermestoides | 0.1 | 0.0 | 0.0 | 0.1 | 2.3 | 0.8 |
| Scolytus multistriatus | 0.8 | 1.8 | 0.8 | 0.1 | 0.1 | 0.5 |
| Hylesinus toranio | 3.2 | 0.0 | 0.1 | 0.1 | 0.1 | 0.5 |
| Hylesinus varius | 0.6 | 0.0 | 0.6 | 0.0 | 0.1 | 0.3 |
| Pteleobius vittatus | 0.0 | 0.0 | 0.8 | 0.1 | 0.0 | 0.2 |
| Scolytus intricatus | 0.2 | 1.2 | 0.3 | 0.0 | 0.2 | 0.2 |
| Scolytus koenigi | 0.2 | 0.2 | 0.2 | 0.3 | 0.1 | 0.2 |
| Anisandrus maiche | 0.4 | 0.7 | 0.2 | 0.0 | 0.0 | 0.2 |
| Total for these species | 97.4 | 98.1 | 99.1 | 99.2 | 99.7 | 99.0 |
Figure 7
Dominance of xylophagous insects in different zones of Gomilshanski lisy National Nature Park
Figure 8
Clusters of sample plots of Gomilshanski lisy National Nature Park by xylophagous insect occurrence
Table 4
Calculation of Sorenson–Chekanovsky index (Csc) for xylophagous complexes in the different sample plots of Gomilshanski lisy National Nature Park
| Group of sample plots (zone) A | Species number in A | Group of sample plots (zone) B | Species number in B | Number of common species for A and B | Csc |
|---|---|---|---|---|---|
| Selective sanitary felling | 22 | Protected | 16 | 12 | 0.63 |
| Selective sanitary felling | 22 | Regulated recreation | 19 | 10 | 0.49 |
| Selective sanitary felling | 22 | Clear felling | 25 | 12 | 0.51 |
| Selective sanitary felling | 22 | Stationary recreation | 22 | 11 | 0.50 |
| Protected | 16 | Regulated recreation | 19 | 11 | 0.63 |
| Protected | 16 | Clear felling | 25 | 13 | 0.63 |
| Protected | 16 | Stationary recreation | 22 | 13 | 0.68 |
| Regulated recreation | 19 | Clear felling | 25 | 12 | 0.55 |
| Regulated recreation | 19 | Stationary recreation | 22 | 12 | 0.59 |
| Clear felling | 25 | Stationary recreation | 22 | 17 | 0.72 |
Figure 9
Competition level between xylophages at the different sample plots of Gomilshanski lisy National Nature Park